FEBRUARY 1st APPLICATION DEADLINE*
*Applications will be accepted past the deadline if space available. For information please contact Stacy Markman, FHL Student Coordinator.
FHL SUMMER A TERM: June 14-July 16 (5 weeks)
Students arrive Sunday, June 13 after 3:00 p.m., depart Friday, July 16 after lunch.
FHL SUMMER B TERM: July 19-August 20 (5 weeks)
Students arrive Sunday, July 18 after 3:00 p.m., depart Friday, August 20 after lunch.
Summer classes held Monday-Saturday:
Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting.
The 5-week courses in summer are intended primarily for graduate students, with the exception of Marine Invertebrate Zoology. Courses may be taken sequentially, i.e., one in each summer session, but not concurrently. Well-qualified undergraduates may be admitted to graduate level courses with the consent of the FHL Director and the faculty involved.
In Summer 2010 FHL is offering one undergraduate-level course and seven graduate-level courses. Each course is 9 credits.
Apply
Costs
Student Information
Spring 2010
Autumn 2010
*Applications will be accepted past due date if space available. For information please contact Stacy Markman, FHL Student Coordinator.
Marine Invertebrate Zoology
Biol 432 (9 undergraduate-level credits)
Session A: June 14 - July 16, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, June 13 after 3 pm, depart Friday, July 16 after lunch.
Dr. Marjorie Wonham and Dr. Noa Shenkar
Comparative biology of marine invertebrate animals, focusing on morphology, natural history, functional biology, life history, and evolutionary relationships. Two daily lectures will provide overviews of the major and many smaller phyla, but the heart of the course comprises study of living animals in the laboratory and fieldwork in the diverse marine habitats surrounding San Juan Island.
Applications are welcome from undergraduate students, post-baccalaureates and graduate students. Prior coursework in invertebrate biology or animal diversity is advisable but not essential.
Enrollment is limited to 20 students.
Photo: Dr. Andrea Ogston
For additional information, contact marwonham@yahoo.com or nshenkar@u.washington.edu
Apply
Costs
Student Information
Session A: June 14 - July 16, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, June 13 after 3 pm, depart Friday, July 16 after lunch.
Dr. Bob Waaland and Dr. Tom Mumford
This course explores the biodiversity of marine algae with emphasis on their role in marine ecosystems. The course will have four major components:
1. Seaweed diversity will be investigated by introducing and practicing the techniques and skills essential for identification of marine macrophytes. Seaweeds from a diversity of habitats (e.g., intertidal, subtidal, sheltered and exposed, eelgrass beds, salt marshes) will be examined through field forays and laboratory studies of seaweed-dominated cool temperate marine communities accessible in the San Juan Archipelago and on the exposed outer coast of Vancouver Island. Collection, preservation and record keeping essential for biodiversity sampling and analysis will be emphasized. Laboratory methods will emphasize the use of essential literature, internet databases, and microscopic examination in order to understand the morphological and reproductive diversity and the details required for identification of diverse seaweed taxa. We plan to include at least two dredging trips on the R/V Centennial for access to the deeper marine flora; we plan to use an underwater ROV to examine seaweed communities in select localities.
2. The functional role of seaweeds in marine ecosystems will be examined through discussion, laboratory and field methods emphasizing the role of seaweeds as primary producers in coastal marine communities. The functional morphology of seaweeds and their interactions (e.g., in their role as primary producers in food webs, their function as structural elements in habitats) with other marine community components will be explored. Lab and field exercises will include introduction to selected analytical gear (e.g., dissolved oxygen meters, nutrient analysis, and data loggers for temperature and light).
Photo by A. Cheng
3. Quantitative analysis of the distributions and abundances of seaweed populations will be investigated with a combination of lectures, field and lab exercises. Emphasis will be placed on study designs, sampling procedures, methods of data analysis and data interpretation. Students will obtain experience with different field methods of sampling seaweeds and with handling and analyzing population and community data. Various approaches for analyzing assemblage or community data will be discussed and supported by computer sessions with relevant software. Practical applications such as the design of monitoring programs at multiple scales will be addressed; prior statistical knowledge is not a prerequisite.
4. Methods for cultivation of seaweeds will be investigated for use at laboratory to commercial scales as tools to elucidate algal life histories, growth rates and development patterns, and physiological responses. The use of mesocosms as experimental systems, and for production of food, chemicals and restoration will be discussed. Special emphasis will be placed on kelp monitoring and restoration techniques in the vicinity of Friday Harbor.
This is a course appropriate for marine biologists, botanists and ecologists as well as oceanographers with interests in marine biodiversity, conservation biology, and coastal ecology with an emphasis on macroalgal primary producers. Graduate students and advanced undergraduates students (juniors, seniors) are encouraged to apply.
The FHL facilities and environment provide the ideal combination of laboratory facilities, housing and a great variety of marine habitat types with high biodiversity representative of cool-temperate marine habitats which are widely distributed throughout temperate regions of the world. Ready access to a diversity of field sites, to small boats, a larger research vessel, labs with seawater aquaria for maintaining specimens and conducting experiments, excellent microscopy facilities, an excellent library and computing facilities with internet access make this an ideal environment for this course.
Enrollment limited to 15 students.
For additional information contact Bob Waaland (jrw@washington.edu) or Tom Mumford (Thomas.Mumford@dnr.wa.gov).
Apply
Costs
Student Information
Session A: June 14 - July 16, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, June 13 after 3 pm, depart Friday, July 16 after lunch.
Dr. Billie Swalla and Dr. Alexandra Eaves
Comparative Invertebrate Embryology will focus on the development of invertebrate phyla with an evolutionary perspective. We will provide extensive hands-on laboratory experience with the fertilization and development of most invertebrate phyla including: Cnidaria, Ctenophora, Platyhelminthes, Nemertea, Brachiopoda, Phoronida, Bryozoa, Mollusca, Polychaeta, Chaetognatha, Echinodermata, Hemichordata and Tunicata.
The lectures will focus on cellular and molecular analysis of evolutionary changes in development as well as reproduction and gametogenesis. We will emphasize morphological processes and discuss similarities and differences in embryos and how they develop. We will read and critique original literature on Comparative Embryology. Several field trips will acquaint students with the rich invertebrate fauna of the San Juan Islands. Class meets Monday-Friday 8-5 and Saturday 8-12. The class is at the graduate student level, but exceptionally qualified undergraduate students are also admitted. We encourage applications from foreign institutions and diverse scientific backgrounds.
Enrollment is limited to 15 students.
For additional information, contact bjswalla@u.washington.edu or alex.eaves@cahs-bc.ca
Apply
Costs
Student Information
Biol 533 (9 credits)
Session A: June 14 - July 16, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, June 13 after 3 pm, depart Friday, July 16 after lunch.
Dr. Emily Carrington, Dr. John Gosline
This course uses an engineering perspective to evaluate the mechanical design of organisms. We will develop an understanding of the diversity of ways organisms construct materials, organize body plans, and interact with their physical environment. We will explore the basic principles of both solid and fluid mechanics, and apply these principles to specific biological examples. The emphasis will be on the design of marine organisms (especially invertebrates and seaweeds), but terrestrial examples will be drawn upon as well.
Photo by L. Coutts
Topics covered will include: basic laws of fluid motion, life in boundary layers, turbulence and mixing, mechanical properties of biological materials, static and dynamic beam theory,
mechanical design for the surf zone, phenotypic plasticity in mechanical design, and biomechanical techniques. This course is quantitative; a basic understanding of calculus and physics is assumed.
Our activities will be focused around two major themes that are current hot topics in the field of biomechanics: 1) Ecomechanics. Organisms must perform within the constraints of their physical environment. How have environmental parameters guided the evolution of organismal form and function and how will future shifts in climate (temperature, water motion, ocean acidification, etc.) affect organismal form and function? 2) Biomaterials. Material scientists are increasingly looking to nature for inspiration in the design of high performance materials, such as the strong underwater adhesives of barnacles, the tough durable tethers of mussels, and the fracture resistance of snail shells. How many other marine biomaterials could be considered “high performing”? To date, relatively few marine biomaterials have been adequately characterized; the richly diverse biota of the San Juan Islands will undoubtedly provide for novel observations. FHL is the ideal setting for this course because of the diversity of marine flora and fauna available and the accessibility of broad range of habitats.
The first three weeks of the course will be devoted to topical lectures, field trips, and laboratory exercises. Problem sets will be assigned to reinforce principles covered in the lectures and labs. Field trips will introduce students to the rich biota of area, as well as the diversity of habitats common to the San Juan Islands. Laboratory exercises will introduce students to state-of-the-art biomechanical techniques and require written lab reports. Topics covered will include: measuring flow in the lab and field, static testing of materials, dynamic testing, beam theory, fracture mechanics, fatigue failure, hydrodynamic forces in a flume, flow visualization, field force measurement (tenacity & hydrodynamic), wave mechanics, measurement and instrumentation (force transducers, field devices, optical methods, etc.).
The final two weeks of the course will be devoted to independent student projects. Students will design their own experiments, develop protocols, gather data, and interpret their results with respect to other related studies. The final products are 1) an oral presentation and 2) a scientific paper.
Enrollment is limited to 15 students.
For additional information contact: ecarring@u.washington.edu or gosline@zoology.ubc.ca
Apply
Costs
Student Information
*Applications will be accepted past due date if space available. For information please contact Stacy Markman, FHL Student Coordinator.
The 5-week courses in summer are intended primarily for graduate students, with the exception of Marine Invertebrate Zoology. Courses may be taken sequentially, i.e., one in each summer session, but not concurrently. Well-qualified undergraduates may be admitted to graduate level courses with the consent of the Director and the faculty involved.
Session B: July 19 - August 20, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 18 after 3 pm, depart Friday, Aug. 20 after lunch.
Dr. Jim Murray, Dr. Shaun Cain
This animation is derived from a confocal image stack of four neurons that have been injected with a fluorescent tracer. The largest cell body is ~100 microns wide. By Dr. Jim Murray.
This 5-week graduate course will focus on learning techniques in neuroethological research such as behavioral recording and analysis, electrophysiology of intact and reduced preparations, voltage clamp and pharmacology, immunohistochemistry, and confocal microscopy of neural structures. Each pair of students will explore a project that helps them to learn the techniques that they need in their own research. Research here will focus on the nudibranch sea slug Tritonia diomedea because it is amenable to neuroethological analysis. Lectures will focus on background necessary to understand fundamental techniques in neurophysiology.
This course is one of very few advanced courses offered worldwide on advanced topics in the neurobiology of behavior. Neuroscience, like molecular biology and genetics, has tended over recent decades to emphasize reductionistic techniques that have been extremely fruitful in illuminating the basic principles of how cells interact to produce nervous systems. But now we find that our ability to collect large data sets of recordings from dozens or hundreds of nerve cells have often outstripped our ability to relate these data back to the behavior of the organism, and ultimately to the ecological context of that behavior. The Tritonia sea slug has served as a model system both in this reductionist approach, but also in a complementary integrative approach to relating the activities of multiple nerve cells to behavior in a natural context. The system is now ripe for an integrative approach to show how animals orient using multimodal sensory cues and how their brains make ecologically-relevant decisions on a cellular level (e.g. should a slug turn right if it smells both food and predator).
Students will be paired-up for 4-week projects, and we will set up a "rig" of electronic equipment for each group. These rigs will include amplifiers of neural activity, analog/digital interfaces to record data onto computers, microscopes to help guide the recording electrodes, and other devices as necessary. We will instruct students in techniques such as intracellular recording, single-cell inactivation, whole nerve recording, and fine-wire recording in freely-moving animals. We will also teach students how to track animal movement in an large tank using cameras computer software, how to analyze video of body movement, and how to correlate movement with neural activity. Students can compare the behavior of sham-operated animals with that of animals whose cells have been inactivated using drugs. Students will also learn to label specific nerve cells using iontophoresis of fluorescent tracers, to immunolabel or use in situ hybridization, and to section and process tissue for confocal microscopy.
Neuroethology has historically been characterized by a focus on comparative, interdisciplinary, integrative, evolutionary, and ecologically-relevant approaches. FHL has traditionally emphasized these approaches is the ideal location for the study of the neuroethology of navigation in Tritonia and other sea slugs.
Links to additional information:
http://gallery.me.com/tritoniadiomedea#100065&bgcolor=black&view=grid
http://depts.washington.edu/fhl/biol533a/
Enrollment limited to 12 students.
Photo: Dr. Jim Murray. The marine slug Tritonia diomedea glides on its foot and explores its world with a touch and taste sensitive oral veil (mustached like anterior tentacles), and it smells food and mates using its posterior head tentacles known as "rhinophores").
For additional information contact: shaun.cain@eou.edu or james.murray@csueastbay.edu
Apply
Costs
Student Information
Biology 533 (9 credits)
Session B: July 19 - August 20, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 18 after 3 pm, depart Friday, Aug. 20 after lunch.
Dr. Drew Harvell, Dr. Carolyn Friedman, and Dr. Steven Roberts
Infectious diseases of marine organisms are on the increase, and yet processes governing host infectivity and pathogen virulence are poorly known, especially for non-commercial marine invertebrates. Indeed, one of the emerging frontiers in ocean research is invertebrate-microbial interactions. Despite these knowledge gaps, the prediction is that diseases will increase in warming oceans and become an ever-present component of near-shore ecological interactions. This course will be a training program in invertebrate-pathogen ecology that will bring together and train the future leaders in this rapidly emerging, multidisciplinary field. The course will 1) survey host-pathogen interaction in the Friday Harbor region, 2) teach diagnostic tools for identifying viral, bacterial, protozoan and fungal infections of invertebrates, 3) teach approaches to examine the invertebrate innate immune response to different pathogens, and finally 4) use these methods to address ecological questions about the distribution of pathogenic interactions, and the experimental effects of temperature and increased acidification on interactions. 
A primary goal of the program is to provide advanced undergraduates, graduate students, and postdoctoral investigators with a broad understanding of host-pathogen interactions as well as the techniques used to study the ecology of marine animals in situ. By bringing together top researchers in host-pathogen interactions, we provide students with a unique opportunity to work side by side with world experts using state-of-the-art tools and technologies. The program also provides a setting for developing and testing new technologies and methods. We also hope it will serve as a research magnet, attracting leading scientists to conduct their own research in a creative teaching and learning environment that catalyzes interactions across the various disciplines associated with Marine Disease Ecology.
In a broader sense, this project will add to our limited understanding of how climate change affects the ecological health of temperate coastal communities. It will also help to address basic information gaps concerning direct biological effects of climate change on susceptible species and the dynamics of parasitism and symbiosis. This understanding is critical to developing realistic management schemes for mitigating impacts of climate change.
Enrollment limited to 15 students.
Photos: Dr. Drew Harvell
For additional information contact: cdh5@cornell.edu, carolynf@u.washington.edu, or sr320@u.washington.edu
Apply
Costs
Student Information
The emphasis will be on functional requirements and constraints for embryos, larvae, and juveniles of marine animals. Topics include maternal investment per ovum, fertilization, parental protection and retention of embryos, extraembryonic nutrition, larval feeding and swimming, functional morphology of embryos and larvae, dispersal, settling, mortality, recruitment, effects of larval nutrition on performance of juveniles, juvenile ecology, and evolutionary transitions between modes of development.D. Grünbaum brings experience with the effects of currents, turbulence, swimming biomechanics and larval behavior on larval distributions. R. Strathmann’s research is on functional constraints on modes of development. The course includes short original research projects, one or occasionally two lectures each day, demonstration of methods, and discussion of papers.
Enrollment limited to 15 students.
Photo: Dr. Richard Strathmann
For additional information contact: random@u.washington.edu or rrstrath@u.washington.edu
Apply
Costs
Student Information
Fish 565 (9 credits)
Session B: July 19 - August 20, 2010 (5 weeks)
Monday-Saturday (Mon-Fri 8 am-5 pm, plus Sat morning 8 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 18 after 3 pm, depart Friday, Aug. 20 after lunch.
Dr. Adam Summers, Dr. Lara Ferry-Graham
The course will use the diverse marine fish community of the San Juan Islands as a tool to explore the relationship between functional morphology and ecology. Students in the course will learn: 1) the evolutionary history and relationships of the major radiations of bony and cartilaginous fishes; 2) basic ecological principles as they relate to fish biology; 3) tools and techniques for collecting fishes; 4) basic morphology of cartilaginous and bony fishes; 5) tools and techniques of functional morphology.
For the first several weeks of the course there will be daily lectures and field trips to familiarize students with the basic tools and animals that they will need for the latter portion of the course. For the second half of the course students will pursue an independent research project. A variety of projects will be suggested but it is also possible to come up with a completely original project based on personal interest. In the past, projects have covered a wide range of topics including ecology, eco-morphology, comparative physiology, comparative morphology and functional morphology. The course will culminate in an oral and written presentation of the results of the research project. This course has historically enjoyed a strong place in the training of functional morphological researchers and the learning goals reflect this.
Enrollment limited to 15 students.
Photos: Dr. Lara Ferry-Graham
For additional information contact: fishguy@u.washington.edu or lfgraham@mlml.calstate.edu
Apply
Costs
Student Information
This non-credit short course begins with a standard check-out dive and includes all the components required to acheieve current scientific diving status with AAUS (the American Academy of Underwater Scientists, see www.aaus.org for more information) and the University of Washington.
The week will include First Aid/CPR and Oxygen for SCUBA emergencies certifications and a full SCUBA rescue course resulting in PADI certification. Other topics that will be covered in either lecture, lab, or class-discussion format are local subtidal animal and algae identification, SCUBA accident management, small-boat handling, and commonly-used methods for gathering biological data underwater.
Applications are welcome from undergraduate students, post-baccalaureates and graduate students from UW or other institutions. Prior marine science experience is recommended but not required. Applicants must be able to show a logbook with a minimum of 20 dives. Students will be required to pass a UW-reviewed physical exam and to have their own SCUBA gear that meets FHL safety standards.
There will be a fee of $950 for this short course, including room and board. Following admission to the course, students will have to pay a $500 non-refundable deposit on or before July 26. Contact Stacy Markman (206-616-0753) with your payment by check or Visa or Mastercard. The remaining $450 is due upon arrival at FHL.
Enrollment limited to 8 students.
Photo: Kevin Turner
For additional information contact:
pema@uw.edu
To apply, students must:
(a) submit the on-line application form (use pull-down menu for Sum B courses), and
(b) fax (206-543-1273) or e-mail the application addendum to Pema Kitaeff.
Application deadline: June 7
During all quarters, graduate students may register for research with the consent of their faculty advisors.
600 Independent Study or Research
700 Master's Thesis
800 Doctoral Dissertation