Friday Harbor Laboratories
last modified July 7, 2015


Summer 2015 Courses, Internships and Workshops


Student Quotes

  • "My experience in FHL was one of the best in my academic background….The course filled out all my expectation about the application of marine conservation sciences in the real world. Also as an Hispanic student I could share my academic and professional experience in Latin America with my classmates and professors."
  • "FHL has an incredible task to educate future marine biologists with a holistic point of view about sciences and that is the reason why I do believe the support for this cause has to continue."
  • "Part of my integral formation as Marine Biologist and Ecologist is due to the course I attended in FHL. It was not only an academic experience but a cultural one. The opportunity to attend one of the courses you offered there was a great experience for an international student."
  • "It got me interested in all sorts of things starting with studies of development of marine invertebrates, and continuing with the nervous system! It exposed me to great joy of exploring the world around us! It changed my life forever! I was able to find great jobs! And, because of FHL I am planning on going to Masters School in marine related science!"


June 15-July 17 (5 weeks), applications review begins Feb. 1 *
Students arrive Sunday, June 14 after 3:00 p.m.
Students depart Friday, July 17 after lunch (served 12:15 - 12:45 p.m.)



MARINE ALGAE (FHL/BIOL 539) - course cancelled as of 2/13/15
SENSORY BIOLOGY AND BEHAVIOR OF FISHES (FHL 528 A) - course cancelled as of 4/13/15


July 20-August 21 (5 weeks), applications review begins Feb. 1*
Students arrive Sunday, July 19 after 3:00 p.m.
Students depart Friday, August 21 after lunch (served 12:15 - 12:45 p.m.)

4. ECOLOGY BETWEEN & BELOW PACIFIC TIDES (with Scientific Diving option) (FHL 568 B)


BLINKS - NSF - BEACON Internship Program: Providing paid research experiences for undergrads, post-bacs or grad students from diverse cultural backgrounds.

Application deadline: March 1, 2015
June 15 - August 8 (8 weeks)


PRACTICAL COMPUTING FOR BIOLOGISTS (AND OTHER SCIENTISTS) (Aug 10 - 21, application deadline February 1)

(Aug. 24 - Sep. 11, application deadline March 15) Anneli Diversity Workshop Cancelled as of 3/18/15

SUMMER WORKSHOP ON THE DYNAMIC BRAIN (Aug 23 - Sep 6, application deadline April 1)

*Students are encouraged to apply for courses as early as possible. Applications will be accepted past the set review date if space is available. For information please contact Stacy Markman, FHL Student Coordinator:

Each course in Summer A term and Summer B term will be 9 credits.

Summer classes are held Monday-Saturday: Mon-Fri 8:30 am-5:00 pm, plus Saturday mornings 8 am-noon, except during the final week of the term when the final Friday is a half-day and there is no Saturday meeting.

Each 5-week course in summer is 9 credits. Courses may be taken sequentially, i.e., one in each summer session, but not concurrently. 400-level courses are undergraduate-level, 500-level courses are graduate-level. Most summer courses are intended primarily for graduate students, with the exception of two undergraduate-level summer courses: Marine Invertebrate Zoology, and Ecology & Conservation of Marine Birds & Mammals. Well-qualified undergraduates may be admitted to a graduate-level course with the consent of the FHL Director and the faculty teaching the course. 400-level courses are considered appropriate for either upper-level undergraduates or graduate students and credits may, in some cases, be applied toward a graduate degree; students should confer with their advisor.

Credits for FHL coursework will be earned through the University of Washington, but applicants to FHL courses do not need to be matriculated at University of Washington. Students from all over the world come to study and conduct research at Friday Harbor Labs.

How do students register for courses at Friday Harbor Labs?

Students must apply and be accepted by Friday Harbor Labs before they can register for FHL courses. Accepted students will be assisted by FHL staff to be registered through University of Washington Professional and Continuing Education (UWPCE). Students, including UW students, may not register themselves for FHL summer courses without assistance from FHL staff.

Additional registration information


Estimated Costs 2015

Frequently Asked Questions - Students

How to travel to Friday Harbor

Student Information

Spring 2015

Autumn 2015

How to request transcripts


Marine Invertebrate Zoology

(FHL/BIOL 432, 9 undergraduate-level credits)
Univeristy of Washington transcript will list "FHL 432: Marine Invertebrate Zoology"

Photo: Mikhail Matz

Summer Term A: June 15 - July 17, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Students arrive Sunday, June 14 after 3:00 p.m., depart Friday, July 17 after lunch.

Dr. Gustav Paulay
University of Florida
Florida Museum of Natural History
paulay at

Dr. Megan Schwartz
University of Washington, Tacoma
School of Interdisciplinary Arts and Sciences
megansc at

Over 90% of the macroscopic species in the marine biosphere are “invertebrates.”

This course takes advantage of the rich marine biota of the Friday Harbor region to teach the principles of invertebrate organization and biodiversity. It emphasizes comparative study of form and function in phylogenetic and ecological contexts.

Photo by Julia Sigwart: Moon Snail

Alternating with two lectures a day, students will study living representatives of most major groups of marine metazoans in the laboratory, and through fieldwork in the diverse marine habitats surrounding San Juan Island. The course reviews the diversity of animal life in an evolutionary and ecological context, focusing on the comparative study of form, function and life history. We will review all animal phyla, and also explore diversity within phyla based on available exemplars.

Biodiversity is one of the most topical subjects in biology, partly because of its accelerating erosion as a result of increasing human pressures and global change. Having a working knowledge of the diversity of life is fundamental to the study of any subject in biology. FHL is arguably the best location in the US for such a course, given the wealth of local diversity and accumulated knowledge of the local fauna built over a century of investigations.

Applications are welcome from advanced undergraduate students, post-baccalaureates and graduate students. Prior coursework in invertebrate biology or animal diversity will be useful; if in doubt, please contact one of the instructors.

Enrollment is limited to 20 students.



Estimated Costs 2015

Student Information

Photo: Dr. Joe Sisneros

Sensory Biology and Behavior of Fishes

(FHL 528 A, 9 credits)

Note: FHL 528 is listed with University of Washington as "Special Topics in Advanced Fish Biology"

Summer Term A

Dr. Joseph Sisneros
University of Washington Departments of Psychology and Biology
sisneros at

Dr. Stephen Kajiura
Florida Atlantic University Department of Biological Sciences
kajiura at

Dr. Paul Forlano
CUNY Brooklyn College Department of Biology
pforlano at

This five-week graduate course will focus on the comparative exploration of the sensory mechanisms that
control behaviors necessary for survival and reproduction in fishes. Through a combination of intensive
lectures and discussions, extensive hands-on laboratory training, one-on-one interactions with course faculty,
lab exercises and student-led projects, students will investigate the sensory biology and behavior of fishes at
the organismal, systems, and cellular levels using modern techniques that include behavioral recordings and
analysis, electrophysiology, neuroanatomy, immunohistochemistry, and brain activation imaging techniques.
Laboratory exercises will include the behavioral analysis of fishes to biologically-relevant stimuli
(electrosensory and lateral line stimuli), fish vocalization recordings and analysis, morphological analysis of
sensory receptor systems and associated brain structures, brain dissection, sectioning and staining, auditory
evoked potential-, electroretinogram and electro-olfactogram recordings, analysis of brain activation patterns
and the functional mapping of neural activity during the expression of behavior via immunohistochemistry for
products of immediate early gene expression.

Enrollment is limited to 15 students.



Estimated Costs 2015

Student Information

Marine Algae


(FHL/BIOL 539, 9 credits)
Univeristy of Washington transcript will list "FHL 539: Marine Algae"

Summer Term A
Using a stadia rod in the intertidal.

Dr. Thomas Mumford
Marine Agronomics, LLC
Olympia, WA
tmumford at

Dr. Brian Wysor
Roger Williams University
Department of Biology, Marine Biology & Environmental Sciences
bwysor at

With nearly 625 species of marine macroalgae (i.e., seaweeds) reported for the region, the San Juan Islands provide one of the most species-rich seaweed assemblages in the world. This charismatic megaflora has played host to generations of phycologists for over 100 years and is the type locality for over 20 species, making it an ideal place to elucidate the principles, methods, and applications of marine macroalgal biodiversity studies. Furthermore, considering the range of human impacts on natural ecosystems and current trends in biodiversity decline from increases in sea surface temperature, ocean acidification or biological invasions, to name but a few, an understanding of the diversity, distribution and ecological role of habitat-structuring marine floras is essential.

In this course, successful students will acquire practical experience in specimen collection, preservation, and databasing, light microscopy, DNA isolation, amplification and sequencing, and computational approaches to phylogeny reconstruction. Extensive field work will be enhanced by lectures as well as laboratory instruction in the observation and interpretation of morphological and anatomical traits that define species and/or higher taxonomic groups. The use of combined approaches will be emphasized to answer basic questions; individual and group projects will use morphological, ecological and molecular data to assess the diversity of algal populations and interpret that diversity in its ecological context. In addition to enhancing technical proficiencies in phycology, all participants will also gain practical experience communicating technical expertise to broader audiences through the production of a public outreach product.

Algae collection aboard the Centennial

At the end of the course, students should be able to use several of the tools now available to identify and classify algae and to critically assess the value of these tools in studies of algal biodiversity and marine benthic ecosystems. Applicants with interests in marine biodiversity, conservation biology, coastal ecology with an emphasis on primary producers, and commercial applications of algae or other diverse scientific backgrounds are encouraged to apply. This is a graduate course, but exceptionally qualified undergraduates will be considered; the course serves as an opportunity to network with dedicated scientists and to develop meaningful research collaborations.

Graduate student members of the Phycological Society of America are eligible to apply for the Hannah T. Croasdale Fellowship, which offers fellowships of up to $1,500 to help defray costs associated with participating in this course. Interested students should also inquire with graduate student organizations and departments at their home institution, which often have awards to help defray costs of participating in field biology courses. Because this course also involves specimen-based research, you may qualify for research awards. Please contact the course instructors to learn more about how the course learning objectives might align with your research goals.

Enrollment limited to 15 students.



Estimated 2015 Costs

Student Information


Photo: Michaelangelo von Dassow

Comparative Invertebrate Embryology

(FHL/BIOL 536, 9 credits)
Univeristy of Washington transcript will list "FHL 536: Comparative Invertebrate Embryology"

Summer Term A: June 15 - July 17, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Students arrive Sunday, June 14 after 3:00 p.m., depart Friday, July 17 after lunch.

Dr. Michelangelo von Dassow
Duke Marine Lab

Dr. Billie Swalla
University of Washington Friday Harbor Laboratories

Comparative Invertebrate Embryology will use hands-on lab experience to introduce students to the great diversity of developmental modes and processes found among marine invertebrates. The course will bridge cell and molecular approaches with ecological and evolutionary approaches to provide an integrated view of animal
development. The course is intended to serve both biologists who wish to understand
diversity in modes of development for ecological and evolutionary studies, and cell and
developmental biologists who wish to broaden their knowledge of embryos beyond the standard model systems.

Photo: Michaelangelo von Dassow

This course provides extensive laboratory experience with fertilization, embryonic and larval development, and metamorphosis of diverse animals. Phyla represented usually include the Porifera, Cnidaria, Ctenophora, Platyhelminthes, Nemertea, Mollusca, Annelida, Brachiopoda, Phoronida, Bryozoa, Echinodermata, Chordata, Chaetognatha, and Arthropoda.

In addition to the basics of invertebrate reproduction and development, lectures will also include analysis of morphogenetic processes, evolutionary changes in development, and functional consequences of different modes of development.

Lab time will be devoted to obtaining, observing and documenting stages of embryogenesis. Lecture and lab practice will also introduce various techniques including (but not limited to) time-lapse microscopy, immunofluorescence, confocal microscopy or in situ hybridization, and biomechanical methods. Field collecting trips to diverse habitats will acquaint students with the environments in which reproduction and development occur and diverse sources of embryos.

Enrollment is limited to 15 students.


Estimated 2015 Costs

Student Information


Ecology and Conservation of Marine Birds and Mammals

(FHL/FISH 492, 9 credits)

Univeristy of Washington transcript will list "FHL 492: Ecology and Conservation of Marine Birds and Mammals"

Photo: Phil Green

Session B: July 20 - August 21, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 19 after 3 pm, depart Friday, Aug. 21 after lunch.

W. Breck Tyler
Institute of Marine Sciences
University of California, Santa Cruz
Long Marine Laboratory

Dr. Eric M. Anderson
Ecological Restoration Program
British Columbia Institute of Technology
Eric_Anderson at

Photo: Phil Green

This intensive, field-based course offers motivated students the opportunity to learn first-hand about the marine birds and mammals of the Salish Sea. Perched at the edge of the San Juan Channel, the Friday Harbor Labs are a great place to learn about these iconic animals and the conservation problems they face, as well as to develop the research skills needed to study them. We welcome applications from undergraduates, post-baccalaureates, and graduate students.

This course emphasizes hands-on learning and makes full use of the Labs’ research boats and facilities. During the first half of the course, lectures, field trips, and lab demonstrations familiarize students with the local fauna, their habitats, and relevant research techniques. Specifically, you will learn: 1) the systematics, morphology, physiology, and ecology of local species; 2) field identification and research techniques for studies of populations, behavior, diet, energetics, and other topics; 3) relationship of tides and other environmental variables to animal distribution and abundance; and 4) the status and conservation of local species.

Recent studies have shown that populations of many marine birds and mammals are declining in the Salish Sea. Therefore, during the latter part of the course, participants conduct independent research on the ecology of local species and communities. Projects may cover a variety of topics and are designed to gather data pertinent to pressing conservation problems. Working in small teams, you will design the project, collect and analyze field data, and then present your results and discuss your findings in a scientific-format paper and presentation.  Examples of recent projects include: effects of tidal currents on Harbor Seal haul out patterns; effects of boat disturbance on marine bird behavior; prey availability and selection of Black Oystercatchers; inter-annual variation in abundance and distribution of auklets, seals, and porpoises; and feeding behavior of Great Blue Herons. Student projects are added to a growing database for the San Juan Island region now being developed by FHL courses and researchers. Your data will help us monitor the status of local species and contribute to future conservation actions.

Enrollment limited to 20.

For more information, contact Breck Tyler: ospr at


Estimated 2015 Costs

Student Information

Ocean Acidification

(FHL 568 A, 9 credits)

Note: FHL 568 is listed with University of Washington as "Special Topics in Advanced Ecology and Biomechanics"

Session B: July 20 - August 21, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 19 after 3 pm, depart Friday, Aug. 21 after lunch.

Photos: Richard Emlet

Dr. Andrew G. Dickson
Marine Physical Laboratory
University of California
adickson at


Dr. Terrie Klinger
School of Marine and Environmental Affairs
University of Washington,
tklinger at


Dr. Jon Havenhand
Dept. Biological & Environmental Sciences
University of Gothenburg
jon.havenhand at

The focus of teaching will be on interactive workshops and discussions of key issues, rather than traditional lectures (although there will be several of those too). Workshops and discussions will focus on critiques of key papers in the literature, which will be selected to present different viewpoints on key topics and engender debate. (As part of the course, you’ll be given guidance on the generic, and essential, skill of critically evaluating a scientific paper, and summarizing its content). In comparison to traditional lectures, this framework provides increased opportunity to anchor basic understanding and analysis methods, provides more flexibility to address key issues in the recent literature, allows us to tailor the content to material that is of relevance to your research, and is quite simply more fun.

Laboratory exercises will focus on establishing sound laboratory practice, which is essential for chemical analyses, but also highly relevant – albeit sometimes neglected – for design and analysis of biological experiments. These exercises will form the foundations on which you’ll build your lab/field project in the second half of the course.

Enrollment limited to 15 students.


Estimated 2015 Costs

Student Information

Fish Swimming:
Kinematics, ecomorphology, behavior, and environmental physiology

(FHL 528 B, 9 credits)
Note: FHL 528 is listed with University of Washington as "Special Topics in Advanced Fish Biology"

Session B: July 20 - August 21, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 19 after 3 pm, depart Friday, Aug. 21 after lunch.

Dr. Paolo Domenici
IAMC- CNR Organismal Biology Lab
Oristano, Italy
paolo.domenici at

Dr. John F. Steffensen
University of Copenhagen
Marine Biological Laboratory
Helsingor, Denmark
JFSteffensen at

Fish swimming is a multidisciplinary area of research that encompasses biomechanics, physiology, evolution, ecology and behavior. Knowledge of fish swimming is relevant both for students interested in mechanisms of locomotion, and those interested in locomotor adaptations to the environment. The course will reflect the multidisciplinary nature of fish swimming. The main subjects treated in the course will be: (1) the kinematics and performance of swimming in fish using various locomotory modes, (2) the ecomorphology of fish locomotion, (3) locomotor strategies, (4) metabolic aspects of fish swimming, (5) the effect of various environmental factors on fish swimming.

Specific lectures will be given on the following topics: Introduction to local fish fauna, introduction to fish hydrodynamics, fish swimming kinematics and biomechanics (steady and unsteady), fish swimming performance (steady and unsteady), scaling of swimming performance, predator-prey encounters, fish functional morphology and swimming, behavioral lateralization in fish swimming, schooling behavior, respiratory physiology, principle of respirometry, ecophysiology of fish swimming, metabolism and exercise physiology, the effect of environmental factors on fish swimming, video analysis techniques, kinematic analysis, circular statistics, and respirometry techniques.

These topics will be treated in lectures and laboratory/field sessions. Students will learn laboratory techniques of video analysis, kinematics, energetics and respirometry. The first half of the course will have an emphasis on lectures and explanations of techniques for studying fish swimming in the laboratory and in the field. In the second half of the course, emphasis will be placed on laboratory and field work. Students will pursue independent research projects. These will be discussed between each student and the instructors. Based on past experience from previous courses taught at FHL, a number of projects will be proposed and rated in terms of their feasibility, their originality and scientific interest. Original projects on fish locomotion, based on the student’s personal background and interest, will also be welcomed. Regular morning meetings will be held in order to discuss various issues such clarifying lecture material, planning logistic matters (fishing, sharing equipment), defining/assigning and updating each project. At the end of the course, students are expected to present the results of their independent projects orally and as a written report in the format of a scientific paper.

The instructors who have designed the course have extensive experience in fish swimming research and teaching. They cover the proposed topics very well. They are internationally-known researchers in the field of fish swimming, and are complementary in terms of expertise. Paolo Domenici is an expert in fish swimming kinematics and behavior, John Steffensen is an expert in fish swimming physiology and energetics. In addition, they taught this course together at FHL in 2002, 2005, 2007, 2009, 2011 and 2013, and they collaborate extensively since 1994. The course proposed has been successful in previous year, as testified by the number of conference presentations (>35) and scientific papers published, in press or in preparation (>15) by the students as a result of their research carried out during the course.

Enrollment is limited to 15 students.


Estimated 2015 Costs

Student Information

Ecology Between & Below Pacific Tides (with Scientific Diving option)

(FHL 568 B, 9 credits)
Note: FHL 568 is listed with University of Washington as "Special Topics in Advanced Ecology and Biomechanics"

Session B: July 20 - August 21, 2015 (5 weeks)
Monday-Saturday (Mon-Fri 8:30 am-5 pm, plus Sat morning 8:30 am-noon, except final week no Saturday meeting)
Arrive Sunday, July 19 after 3 pm, depart Friday, Aug. 21 after lunch.

Photo: Robin Elahi

Dr. Aaron Galloway
Washington State University, School of the Environment
aaron.galloway at

Tiffany Stephens
University of Otago, New Zealand
tiffanybot at

Pema Kitaeff
University of Washington
Friday Harbor Laboratories
pema at

Dr. Megan Dethier
University of Washington
Friday Harbor Laboratories
mdethier at

Dr. David Duggins
University of Washington
Friday Harbor Laboratories
dduggins at

Marine ecology remains a vibrant field relevant to a variety of modern environmental issues at local and global scales. For example, near-shore communities are being influenced by the modification of shorelines, excessive nutrient and sediment inputs, species invasions, and seawater warming and acidification. A solid background in the essentials of marine ecology and in the fundamental techniques used to study ecological questions in the marine realm is needed by students entering any field that involves ecological or environmental issues in the ocean.

This unique marine ecology course will focus on both intertidal and subtidal theory and methods, bringing together experts from multiple fields of marine ecology to guide students through the course. The course has two tracks, one that includes AAUS certification for SCUBA divers interested in scientific diving, and the other for non-divers interested in field ecology and laboratory experiments. Students in the diving track will spend the first two weeks honing their rescue, navigation, and data collection techniques underwater. Training will emphasize safe, efficient methods to conduct subtidal fieldwork. For the non-diving track, in the first two weeks students will explore the range of local rocky and soft-sediment intertidal habitats, practice sampling and survey methods in the intertidal zone, and run a variety of laboratory experiments. Interested students may attend some of the diving-related lectures, such as CPR certification.

Together, all students will learn marine life identification and work in the range of marine ecosystems present in the San Juan Islands. Instructors will present the fundamentals of marine ecology, including productivity and limiting factors, diversity, species interactions, disturbance and recruitment dynamics. Students will identify relevant research question of their own choosing or suggested by instructors. The last three weeks of the course will be spent developing methods, collecting and analyzing data, and preparing research papers and presentations.

If applying for the Scientific Diving option, please submit a general FHL application form and then download and complete the Scientific Diving Application Addendum and submit via email to Pema Kitaeff as a PDF file. There will be an additional fee for the Scientific Diving option.

Enrollment is limited to 20 students.


Estimated 2015 Costs

Student Information

Practical Computing for Biologists (and other scientists)

August 10 - August 22 (2 weeks), applications due Feb. 1*
Workshop participants arrive Sunday, Aug. 9 after 3:00 p.m. (program includes dinner on Sunday, Aug. 9 through breakfast on Saturday, Aug. 22)

Practicial Computing Workshop Participants should arrive at FHL on Sunday, Aug. 9, after 3:00 p.m.

First meal for workshop participants is dinner served Sunday, Aug. 9.

1st class Monday, Aug. 10, 8:30 a.m.

Classes held Mon-Sat 8:30 a.m.-5:00 p.m., plus Sunday, Aug. 16 partial day (time TBD) and possible evening sessions.
Classes end Friday, Aug. 21. Students should plan to depart FHL on Saturday morning, Aug. 22.


Dr. Steven Haddock
Monterey Bay Aquarium Research Institute and U.C. Santa Cruz
haddock at

Dr. Casey Dunn
Brown University
casey_dunn at

This course covers some of the simple but powerful skills that all scientists should know in a world of increasingly complex analyses. It is based on the book Practical Computing for Biologists, from Sinauer Associates, written by the instructors. This is not strictly a bioinformatics or programming course, although the lessons are applicable to molecular data. The skills are applicable to ecology, neurobiology, zoology — any subdiscipline where gathering and analyzing moderate to large data sets are required. The specific sections include working with text files, command-line operations, scripting and Python programming, creating scientific graphics, working with servers (remote login, software installation), databases, version control and a bit about the web and data-acquisition hardware. Skills are immediately applicable, and the most common comment we get from class participants and readers of the book is "This would have save me hours just last week," or "I wish I knew about this when I was in grad school!" The target audience for the course is anyone with data: grad students, post-docs, technicians, and faculty. There are no hard pre-requisites, but participants should bring their own computer — preferably one that they use in their daily work.

Applicants for this workshop will need to submit two forms:
1) the FHL Application Form (link here), and
2) the Practical Computing for Biologists Application Form (link here).

Note: For this workshop we are *not* requiring transcripts or letters of recommendation. The personal statement can be the one that applicants enter on the Practical Computing web form.


Estimated 2015 Costs

Student Information

Annelid Diversity and Evolution Workshop


August 24 - September 12 (3 weeks), application due date extended (originally March 15*) APPLICATIONS STILL BEING ACCPETED

Application Requirements: Please submit an FHL Application Form, a CV, a personal statement (statement of interest) and one or two letters of recommmendation. (Transcripts are *not* required to apply for this non-credit workshop.)

FHL Application Form:

The CV and statement may be submitted by email from the applicant to: The letter(s) or recommendation should be submitted via email directly from the writer to:

Workshop participants arrive Sunday, Aug. 23 after 3:00 p.m. (program includes dinner on Sunday, Aug. 23 and breakfast on Saturday, Sept. 12). Cost: $1800 (includes workshop fee, lab fee, housing and 20 meals per week for three weeks). All fees to be paid to Friday Harbor Labs.

Dr. Ken Halanych
Department of Biological Sciences
Auburn University
ken at

Dr. Anja Schulze
Department of Marine Biology
Department of Oceanography
Texas A&M University at Galveston
schulzea at

Dr. Damhnait McHugh
Division of Natural Sciences & Mathematics
Colgate University
dmchugh at

Dr. Frank Edward (Andy) Anderson
Department of Zoology
Southern Illinois University
feander at

Annelids comprise >16000 recognized species that occupy a wide range of marine, freshwater and terrestrial environments. Many members of this ancient group are ecologically and economically important as ecosystem engineers, as the basis of commercial enterprises, as indicators of environmental health, as invasive or pest species and as being arguably the most abundant metazoans in the deep sea, which covers ∼60% of the planet. As one of the few segmented phyla, annelids are key to understanding the evolution of bilaterian body plans.

Molecular tools have improved our insight into the composition of the group and some relationships within it. The current grant supporting the workshop has been addressing many fundamental questions about annelid phylogeny remain unresolved: What are the major clades within Annelida? Which annelid lineages are basal? What are the sister taxa to Sipuncula and Clitellata? What are the evolutionary transitions to a terrestrial lifestyle in clitellates? Are myzostomids within annelids (sensu stricto)? Equally compelling are questions about the tips of the annelid tree, such as placement of monospecific or highly derived taxa, and whether so-called “cosmopolitan” species in fact represent cryptic species complexes. Major morphologically defined annelid groups are not supported by molecular data; our efforts are leading to a sound phylogenetic framework based on a range of genetic and genomic data. This will provide a better understanding of morphological evolution within the group and across bilaterians.

The course will have a focus on annelid diversity at the organismal level by integrating three key areas; functional morphology, phylogenetic theory, and genomic resources. We will explore these topics on time scales that span the history of the group on the planet. For example, we will use phylogenetics and genomic data to examine the originals annelids and placement of taxa such as Echiura, Sipuncula and Clitellata within the annelid radiation. On the other extreme we will explore the genetic connectivity of present data species, for example, in the Northern Pacific (USA northeastern Pacific to Japan). These discussions will be rooted in a working knowledge of organismal form and function. How have these animals changed during the time scales under consideration? Although the focus will be largely on evolution and biodiversity questions, we will also touch on other topics to cover the breadth of annelid biology including ecology, feeding, reproduction and development. The course will be predominately marine, but terrestrial and aquatic forms will also be covered in both lecture and fieldwork.

Enrollment limited to 15 students.

Student Information


Summer Workshop on the Dynamic Brain


Co-hosted by the Allen Institute for Brain Science and the Computational Neuroscience Program at the University of Washington and directed by Drs. Christof Koch and Adrienne Fairhall.

August 23 - September 6, 2015
Application deadline: April 1

This workshop is being held on the FHL campus but current information and the application process is being managed through this website:

Questions? Please contact: dynamicbrain at


Independent Study for UW Graduate Students

During all quarters, graduate students may register for research with the consent of their faculty advisors.

600 Independent Study or Research
700 Master's Thesis
800 Doctoral Dissertation