(in progress)
With the help of:
D. Albach, B. Bremer, P. Cantino, C. dePamphilis, P. Garnock-Jones, R. Harley, L. McDade, E. Norman, B. Oxelman, J. Reveal, R. Scotland, J. Smith, E. Wallander, A. Weber, A. Wolfe, N. Young, M. Zjhra, and others
[estimated # species in Lamiales = 22,000]
The goal of this project is to produce a working infraordinal classification of the Lamiales to genus with information on distribution and species richness. All recognized taxa will be clades; adherence to Linnaean ranks is optional. Synonymy is very incomplete (comprehensive synonymy is not a goal of the project, but could be incorporated). Although I anticipate producing a publishable version of this classification at a future date, my near-term goal is to produce a web-accessible version, which will be available to the public and which will be updated regularly through input from systematists familiar with taxa within the Lamiales. For further information on the project and to provide information for future versions, please contact R. Olmstead via email at olmstead@u.washington.edu, or by regular mail at: Department of Biology, Box 355325, University of Washington, Seattle WA 98195, USA.
Families treated:
Acanthaceae
Avicenniaceae
Bignoniaceae
Byblidaceae
Calceolariaceae
Carlemanniaceae
Gesneriaceae
Lamiaceae
Lentibulariaceae
Linderniaceae
Martyniaceae
Oleaceae
Orobanchaceae
Paulowniaceae
Pedaliaceae
Phyrmaceae
Plocospermataceae
Schlegeliaceae
Scrophulariaceae
Stilbaceae
Tetrachondraceae
Verbenaceae
Veronicaceae
Genera of Lamiales unassigned to family
Acanthaceae (~201/3510) Durande, Notions El m. Bot.: 265. 1782, nom. cons. Synopsis compiled by R. Scotland & K. Vollesen (Kew Bull. 55: 513-589. 2000); probably should include Avicenniaceae.
Nelsonioideae (7/ ) Lindl. ex Pfeiff., Nomencl. Bot. 1(1): 10. 8 Dec 1871
Anisosepalum
E. Hossain (3) C. Africa
Elytraria
Michx. (17) warm & trop. NW & OW
Gynocraterium
Bremek. (1) trop. S. Amer.
Nelsonia
R. Br. (1) trop. OW
Ophiorrhiziphyllon
Kurz (5) SE Asia
Saintpauliopsis
Staner (1) trop. Africa
Staurogyne
Wall. (140) trop. OW & NW
Thunbergioideae (including Mendoncia) (5/ )
Kostel., Allg. Med.-Pharm. Fl. 3: 923. Apr-Dec 1834
Anomacanthus
R.D. Good (1) Zaire, Angola
Mendoncia
Vell ex Vand. (60) trop. Amer. & Africa, Madag.
Meyenia
Nees (1) India, Sri Lanka
Pseudocalyx
Radlk. (4) trop. Africa, Madag.
Thunbergia
Retz.
(90) trop. OW
Acanthoideae (189/ ) Link, Handbuch 1:500. Jan-Aug 1829
Acanthopsis
Harv. (7) S. Africa
Acanthus
L. (30) warm & trop. OW
Achyrocalyx
Benoist (4) Madag.
Aphelandra
R. Br. (175) Trop. Amer.
Blepharis
Juss. (8) OW trop., Medit., S. Africa
Crossandra
Salisb. (50) trop. Africa & Asia, Madag., Arabian Penin.
Crossandrella
C.B. Clarke (2) trop. Africa
Cynarospermum
Vollesen
Cyphacanthus
Leonard (1) Columbia
Encephalosphaera
Lindau (2) trop. S. Amer.
Geissomeria
Lindl. (15) Mex., C. Amer., trop. S. Amer.
Holographis
Nees (10) Arid Mexico
Neriacanthus
Benth. (4) trop. Amer.
Orophochilus
Lindau (1) Peru
Rhombochlamys
Lindau (2) Columbia
Salpixantha
Hook. (1) Jamaica
Sclerochiton
Harv. 12) trop. & S. Africa
Stenandrium
Nees (25) warm S. Amer.
Streptosiphon
Mildbr.(1) trop. E. Africa
Strobilacanthus
Griseb. (1) Panama
Xantheranthemum
Lindau (1) Peru
Ruellieae
Ruelliinae
(48/ )
Acanthopale
C.B. Clarke (150) OW trop.
Aechmanthera
Nees(3) Himal.
Apassalus
Kobuski (3) SE US, W.I.
Beniocanthus
Heine & A. Raynal (2) Madag.
Blechum
P. Brown (6) trop. Amer.
Bravaisia
DC. (3) trop. Amer.
Brillantaisia
P. Beauv. (9) trop. Africa, Madag.
Brunoniella
Bremek.(6) New Guinea, Australia
Calacanthus
T. Anderson ex Benth. (1) Indomal.
Clarkeasia
J.R.I. Wood
Dischistocalyx
T.Anderson ex Benth. (20) Africa
Duosperma
Dayton(12) trop. & S. Africa
Dyschoriste
Nees (65) warm & trop. NW & OW
Echinacanthus
Nees(4) Himal., China
Epiclastopelma
Lindau (2) trop. Africa
Eranthemum
L. (30 trop. Asia
Eremomastax
Lindau
Eusiphon
Benoist (3) Madag.
Hemigraphis
Nees (90) Asia to New Caledonia
Heteradelphia
Lindau (1) W. Africa
Hygrophila
R. Br. (25) trop. OW & NW
Ionacanthus
Beniost (1) Madag.
Kosmosiphon
Lindau (1) trop. W. Africa
Leptosiphonium
F. Muell. (10) Papauasia
Louteridium
S. Watson (6) Mex. & C. Amer.
Lychniothyrsus
Lindau (5) Brazil
Mellera
S. Moore (5) warm Africa
Mimulopsis
Schweinf. (3) trop. Africa, Madag.
Pararuellia
Bremek. (5) Malaysia
Petalidium
Nees (35) trop. & S. Africa, Himal., India
Phaulopsis
Willd. (20) Afra. Mascarenes, India
Physacanthus
Benth. (5) trop. Africa
Polylychnis
Bremek. (2) NE S. Amer.
Pseudoruellia
Benoist (1) Madag.
Ruellia
L. (150) temp. & warm to trop. Amer.
Ruelliospis
C.B. Clarke (3) trop. & S. Africa
Sanchezia
Ruiz & Pav.(20) trop. Amer.
Satanocrater
Schweinf. (4) trop. Africa
Sautiera
Decne. (1) Timor
Spirostigma
Nees (1) Brazil
Stenosiphonium
Nees (3) India, Sri Lanka
Stenothyrsus
C.B. Clarke (1) Malaya
Strobilanthes
Blume (250) trop. Asia
Strobilanthopsis
S. Moore (5) trop. Africa
Suessenguthia
Merm. (1) Bolivia
Trichanthera
Kunth (2) N. S. Amer.
Trichosanchezia
Mildbr. (1) trop. Peru
Zygoruellia
Baill. (1) Madag.
Andrographis
Wall. ex Nees (20) trop. Asia
Cystacanthus
T. Anderson (10) SE Asia
Diotacanthus
Benth. (2) Indomal.
Graphandra
J.B. Imlay (1) Thailand
Gymnostachyum
Nees (30) India to C Malaysia.
Haplanthodes
Kuntze (4) India
Indoneesiella
Sreem. (2) India
Phlogacanthus
Nees (15) Indomal.
Justiciinae (103/ )
Afrofittonia
Lindau (1) trop. W. Africa
Ambongia
Beniost (1) Madag.
Ancistranthus
Lindau (1) Cuba
Angkalanthus
Balf.f. (2) Socotra, Transvaal
Anisacanthus
Nees (8) SW US, Mex.
Anisotes
Nees (19) trop. Africa, Madag.
Anthacanthus
Nees (?) Unknown
Aphanosperma
T.F. Daniel (1) NW Mex.
Ascotheca
Heine (1) trop. W. Africa
Asystasia
Blume (70) OW trop.
Ballochia
Balf.f. (3) Socotra
Brachystephanus
Nees (10) trop. Africa, Madag.
Calycacanthus
K. Schum. (1) New Guinea
Carlowrightia
A.Gray (23) SW US, Mex, C. Amer.
Celerina
Benoist (1) Madag.
Centrilla
Lindau (?) Unknown
Cephalacanthus
Lindau (1) Peru
Chalarothyrsus
Lindau (1) Mex.
Chamaeranthemum
Nees(4) trop. Amer.
Chileranthemum
Oerst. (2) Mex.
Chlamydocardia
Lindau (4) trop. W. Africa
Chlamydostachya
Mildbr. (1) trop. E. Africa
Chorisochroa
Vollese
Clinacanthus
Nees (2) S. China to Malaysia
Clistax
Mart. (2) Brazil
Codonacanthus
Nees (2) NE India, S. China, Japan
Conocalyx
Benoist (1) Madag.
Cosmianthemum
Bremek. (8) W. Borneo
Cyclacanthus
S. Moore (2) SE. Asia
Cylindrosolenium
Lindau (1) Peru
Danguya
Beniost (1) Madag.
Dasytropis
Urb. (1) E. Cuba
Dichazothece
Lindau (1) E. Brasil
Dicladanthera
F. Muell. (2) W. Australia
Dicliptera
Juss. (150) warm & trop. NW & OW
Ecbolium
Kurz (19) trop. & S. Africa, Madag., Socotra, India
Filetia
Miq. (8) Sumatra, Indomal.
Fittonia
Coem. (2) Peru
Forcipella
Baill. (5) Madag.
Glossochilus
Nees (2) S. Africa
Graptophyllum
Nees (10) Australia, SW Parcific Isl.
Gypsacanthus
Lott, Jarmillo & Rzed. (1) Mex.
Harpochilus
Nees (3) Brasil
Henrya
Nees (2) C. Amer.
Herpetacanthus
Nees (10) Panama to Brasil
Hoverdenia
Nees (1) Mex.
Hypoestes
Sol. Ex R.Br. (40) OW trop.
Ichtyostoma
Hedren and Vollesen
Isoglossa
Oerst. (50) OW trop. & Arabian Penin.
Isotheca
Turrill (1) Trinidad
Jadunia
Lindau (1) New Guinea
Juruasia
Lindau (2) Brazil
Justicia
L. (600) trop. & warm temp. NW
Kalbreyeriella
Lindau (3) Panama, Columbia
Linariantha
B.L. Burtt & Ros.M.Sm. (1) Borneo
Mackaya
Harv. (15) Africa
Marcania
J.B. Imlay (1) Thailand
Megalochlamys
Lindau (3) SW Africa
Megalostoma
Leonard (1) C. Amer.
Megaskepasma
Lindau (1) Venezula
Mellitacanthus
S. Moore (1) Madag.
Metarungia
Baden (3) trop. & S. Africa
Mexacanthus
T.F. Daniel (1) Mex.
Mirandea
Rzed. (2) Mex.
Monechma
Hochst. (60) Africa, India
Monothecium
Hochst. (3) trop. Africa, S. India
Odontonema
Nees 936) Mex., C. & S. Amer., W. I.
Oplonia
Raf. (19) trop. Amer., Africa (extinct?), Madag.
Oreacanthus
Benth. (4) C. Africa
Pachystachys
Nees (12) trop. Amer.
Pelecostemon
Leonard (1) Columbia
Peristrophe
Nees (15) trop. OW
Phialacanthus
Benth. (5) Himal. to Malay Penin.
Podorungia
Baill. (1) Madag.
Poikilacanthus
Lindau (6) trop. Amer.
Populina
Baill. (2) Madag.
Pranceacanthus
Wassh. (1) Brazil
Pseuderanthemum
Radlk. (60) trop. OW & NW
Pseudodicliptera
Benoist (2) Madag.
Psilanthele
Lindau (1) Ecuador
Ptyssiglottis
T. Anderson (60) Indomal.
Pulchranthus V.M. Baum, Reveal and Nowicke (4) trop. S.
Amer.
Razisea
Oerst. (3) C. Amer.
Rhinacanthus
Nees (15) trop. OW
Ritonia
Benoist (3) Madag.
Rungia
Nees (50) trop. OW
Ruspolia
Lindau (4) trop. Africa
Ruttya
Harv. (3) trop. & S. Africa, Arabian Penin.
Samuelessonia
Urb. & Ekman (1) Hispaniola
Sapphoa
Urb. (2) Cuba
Schaueria
Nees (8) Brazil
Sebastiano-Schaueria
Nees (1) Brazil
Spathacanthus
Baill. (5) C. Amer.
Sphinctacanthus
Benth. (1) NE India to Burma
Stenostephanus
Nees (6) S. Amer.
Streblacanthus
Kuntze (6) C. Amer.
Tessmanniacanthus
Mildbr. (1) trop. Peru
Tetramerium
Nees (28) C. Amer.
Thysanostigma
J.B. Imlay (2) Thailand, Malayan Penin.
Trichaulax
Vollesen
Trichocalyx
Balf.f. (2) Socotra
Xerothamnella
C.T. White (2) E. Australia
Yeatesia
Small (3) SE US, NE Mex.
Barleria
L. (250) OW & NW trop.
Barleriola
Oerst. (6) W. I.
Borneacanthus
Bremek. (6) Borneo
Boutonia
DC. (1) Madag.
Chroesthes
Beniost (3) China, SE Asia
Crabbea
Harv. (12) trop. & S. Africa
Hulemacanthus
S. Moore (1) New Guinea
Lepidagathis
Willd. (100) warm & trop. NW & OW
Lophostachys
Pohl. (15) trop. Amer.
Acanthostelma
Bidgood and Brummitt (1) Somalia
Acanthura
Lindau (1) Brazil
Aphelandrella
Mildbr. (1) Peru
Camarotea
Scott Elliot (1) Madag.
Chlamydacanthus
Lindau (4) trop. E. Africa, Madag.
Dolichostachys
Benoist (1) Madag.
Golaea
Chiov. (1) Somalia
Idiacanthus
Leonard
Kudoacanthus
Hosok. (1) taiwan
Lankesteria
Lindl. (7) trop. Africa, Madag.
Lasiocladus
Bojer ex Nees (5) Madag.
Leandriella
Benoist (1) Madag.
Morsacanthus
Rizzini (1) Brazil
Neuracanthus Nees (20) trop. Africa, Arabian Penin. to
India
Perenideboles
Ram. Goyena (1) Nicaragua
Pericalypta
Beniost (1) Madag.
Sphacanthus
Benoist (2) Madag.
Vavara
Benoist (1) Madag.
Vindasia
Benoist (1) Madag.
Whitfieldia
Hook. (10) trop Africa
Avicenniaceae 1/4-7 Endl., Ench. Bot.: 314. 15-21 Aug 1841 - clearly
belongs within Acanthaceae (Olmstead unpubl., McDade, unpubl.)
Avicennia (4-7) warm cosmop
Bignoniaceae (82/~809) Durande, Notions Elm. Bot.: 265. 1782, nom. cons. - Synoptical classification by tribe following Gentry
1980, 1992; Spangler & Olmstead 1999, Zjhra et al. 2003, Lohman 2005, Grose
& Olmstead, in prep.; Olmstead et al. unpubl.
Bignonieae Dumort., Anal. Fam. Pl.: 23. 1829 (treatment following L. Lohmann, 2005) (21/367)
Adenocalymma Mart. ex Meisn. (78) trop Am
Amphilophium Kunth (41) trop Am
Anemopaegma Mart. ex Meisn. (41) trop Am
Arrabidaea DC. (69) trop Am
Bignonia L. (28) trop Am (including SE USA)
Callichlamys Miq. (1) trop Am
Ceratophytum Pittier (= Paragonia Bureau)
Clytostoma Miers ex Bureau (= Bignonia L.)
Cuspidaria DC. (16) trop Am
Cydista Miers (= Bignonia L.)
Distictella Kuntze (= Amphilophium Kunth)
Distictis Mart. ex. Meisn. (= Amphilophium Kunth)
Dolichandra Cham. (9) trop Am
Fridericia (= Arrabidaea DC.)
Gardnerodoxa Sandwith (= Neojobertia Baill.)
Glaziova Bureau (= Amphilophium Kunth)
Haplolophium Cham. (= Amphilophium Kunth)
Leucocalantha Barb. Rodr. (= Pachyptera DC. ex Meisn.)
Lundia DC. (13) trop Am
Macfadyena A. DC.( = Dolichandra Cham.)
Macranthisiphon Bureau ex K. Schum. (= Bignonia L.)
Manaosella J.C. Gomes (1) Bolivia and Brazil
Mansoa DC. (11) trop Am
Martinella Baill. (2) trop Am
Melloa Bureau (= Dolichandra Cham.)
Memora Miers (= Adenocalymma Mart. ex Meisn.)
Mussatia Bureau ex Baill. (= Bignonia L.)
Neojobertia Baill. (2) Eastern Brazil
Pachyptera DC. ex Meisn. (4) trop Am
Parabignonia Bureau ex K. Schum. (= Dolichandra Cham.)
Paragonia Bureau (11) trop Am
Perianthomega Bureau ex Baill. (1) SE Brazil, NE Parag. and SE Bolivia
Periarrabidaea A. Samp. (= Paragonia Bureau)
Phryganocydia Mart. ex Bureau (= Bignonia L.)
Piriadacus Pichon (= Arrabidaea DC.)
Pithecoctenium Mart. ex Meisn. (= Amphilophium Kunth)
Pleonotoma Miers (15) trop Am
Potamoganos Sandwith (= Bignonia L.)
Pseudocatalpa A. H. Gentry (= Paragonia Bureau)
Pyrostegia C. Presl (3) trop S Am
Roentgenia Urb. (= Bignonia L.)
Saritaea Dugand (= Bignonia L.)
Spathicalyx J. C. Gomes (= Paragonia Bureau)
Sphingiphila A. H. Gentry (1) Paraguay (tentatively
placed here by LL)
Stizophyllum Miers (3) trop Am
Tanaecium Sw (= Bignonia L.)
Tynanthus Miers (17) trop Am
Urbanolophium Melch. (= Amphilophium Kunth)
Catalpeae
DC. ex Meisn. (3/12)
Catalpa Scop. (10) E Asia, N Am
Chilopsis D. Don (1) SW US, Mex
Chitalpa (= Catalpa X Chilopsis)
Macrocatalpa (Griseb.) Britton (1) Cuba (~Catalpa)
Oroxyleae
Gentry 1980 (4/6)
Hieris
Steenis (1) Penang
Millingtonia
L.f. (1) SE Asia, Malaysia
Nyctocalos
Teijsm. & Binn. (3) Indomalaysia
Oroxylum
Vent. (1) Indomalaysia
Old
World Spathodea/Coleeae clade (19/~138)
Coleeae
Bojer 1837 (5/50)
Colea Bojer ex Meisn. (21) Madagascar, Mascarenes
Ophiocolea H, Perrier (5) Madagascar, Comores
Phyllarthron
DC. (15) Madagascar, Comores
Phylloctenium Baill. (2) Madagascar
Rhodocolea
Baill. (7) Madagascar
Old World Spathodea/Coleeae clade unassigned (14/88)
Catophractes D. Don (1) tropical & S Africa
Dinklageodoxa Heine & Sandwith (1) Liberia
Dolichandrone (Fenzl) Seem. (10) E Afr, Indomal, trop.
Austral, New Caledonia
Fernandoa Welw. ex Seem. (14) Afr, Mad, S China, SE
Asia to Sumatra
Haplophragma Dop. (=Fernandoa)
Heterophragma DC. (2) India, SW Asia
Kigelia DC. (2) tropical Africa
Markhamia Seem. ex Baill. (10) trop Afr & SE Asia
Mayodendron Kurz (= Radermachera)
Newbouldia Seem. ex Bureau (1) tropical W Africa
Perichlaena Baill. (1) Madagascar
Radermachera Zoll. & Moritzi. (17) SE Asia,
Malaysia
Rhigozum Burch. (7) trop & S Afr, Ethiop, Somal,
Madag
Spathodea P. Beauv. (1) tropical Africa
Stereospermum Cham. (20) trop Afr & Asia
Tecomella Seem. (1) Arabia to W India
New World - Tabebuia /Crescentieae clade (Digitifolieae) (14/~154)
Crescentieae
G. Don, Gen. Hist. 4: 216, 232. 1837-8 Apr 1838 (3/33)
Amphitecna Miers (18) tropical Am
Crescentia L. (6) tropical Am
Enallagma (Miers) Baill. (= Amphitecna)
Parmentiera DC. (9) C Am, NW Columbia
New World - Tabebuia /Crescentieae clade unassigned (11/~121)
Cybistax Mart. ex Meisn. (1)
Ekmanianthe Urb. (2) Cuba, Hispaniola
Handroanthus (34) Neotrop.
Godmania Hemsl. (2) tropical Am
Paratecoma Kuhlm. (1) Brazil
Romeroa Dugand (1) Colombia
Roseodendron (2) Neotrop.
Sparattosperma Mart. Ex Meisn. (2) trop. S. Am.
Spirotecoma Baill. ex Dalla Torre & Harms (4) Cuba,
Hispaniola
Tabebuia Gomes ex DC. (~70) tropical Am
Zeyheria
Mart. (2) Brazil, Bolivia
Tecomeae Endl., Gen. Pl.: 711. Jan 1839 (10/51)
Campsidium Seem. (1) Chile, Argentina
Campsis Lour. (2) E Asia, E US
Deplanchea
Vieill. (5) Malaysia, Aust, New Cal
Incarvillea Juss. (16) Himalaya, C to E Asia
Lamiodendron
Steenis (1) New Guinea
Pandorea (Endl.) Spach (6) E Malay, Austr, New Cal
Podranea Sprague (1) tropical to S Africa
Tecoma Juss. (12) tropical Am (Andes) to Ariz
Tecomanthe Baill. (5) Malaysia, Austr, NZ
Tecomaria (Endl.) Spach (2) E & S Africa
Tourrettieae G. Don, Gen. Hist. 4: 215, 231. 1837-8
Apr 1838 (2/4)
Eccremocarpus Ruiz & Pav. (3) Colombia to Chile
Tourrettia
Foug. (1) Andes to Mexico
Jacarandeae Seem., Ann. Mag. Nat. Hist., ser. 3, 10:
31. Jul 1862 (2/52)
Digomphia Benth. (3) Guayana highland
Jacaranda Juss. (49) Guatemala/Antilles to Argentina
Bignoniaceae unassigned to more inclusive clade (7/25)
Argylia D. Don (12) Peru, Chile, Argentina
Astianthus D. Don (1) C Am
Barnettia
(=Santisukia)
Delostoma D. Don (4) Andes
Neosepicaea Diels (4) N Guinea, Queensl,, Moluccas
Pajanelia DC. (1) Indomalaysia
Pauldopia Steenis (1) NE India to SE Asia
Santisukia
Brummitt (2) Thailand
Byblidaceae (Engl. & Gilg) Domin, Acta Bot. Bohem. 1:3. 1922,
nom. cons. (1/2)
Byblis (2) Aus, New Guinea
Calceolariaceae (3/275) (G. Don) R. Olmstead 1999 (Olmstead et al.
1999) (see Molau 1988)
Calceolaria
L. (240-270 sensu Molau) C to S Am Porodittia G. Don (1) Peru
Jovellana
Ruiz & Pav. (6) NZ, Chile
Carlemanniaceae Airy Shaw (1964) (APG II, 2002) (2/5)
Carlemannia Benth (3) Indomal
Silvianthus Hook.f. (2) India to SE Asia
Gesneriaceae
(~135-140/>3000) synopsis compiled by A. Weber, provisional state 1999 (classification after Wiehler & Burtt 1995; modified
following J. Smith)
Achimenes Pers. (24), mainly Mexico and Central America
Anodiscus Benth. (1), Peru
Bellonia L. (2), Cuba and Hispaniola
Capanea Planch. (11), Peru to Guatemala
Diastema Benth. (>20), Bolivia to Mexico, Venezuala,
Colombia, Ecuador, Peru
Eucodonia Hanst. (2), C and S Mexico
Gloxinia L'Hr.(15), central America and trop. S
America
Goyazia Taub. (2), C Brazil
Heppiella Regel (4), Peru, W Venezuela
Koellikeria Regel (1), C and trop. S America
Kohleria Regel (>20) , Peru to Mexico, Venezuela,
Trinidad, Guianas
Lembocarpus Leeuwenb. (1), Surinam and French Guiana
Monopyle Benth. (>27), Bolivia to Guatemala
Moussonia Regel (11), Mexico to Panama
Niphaea Lindl. (5), Mexico, Guatemala, Venezuela, Peru
Pearcea Regel (incl. Parakohleria Wiehler) (17),
eastern slopes of the Andes (Colombia, Ecuador, Peru, Bolivia), 1 sp. also on
the western slopes (Ecuador)
Phinaea Benth. (>16), Mexico to Peru, Venezuela,
Cuba, Haiti
Smithiantha Kuntze (5), C & N Mexico, Guatemala
Solenophora Benth. (> 20), central America
Paliavana Vand. (8), SE Brazil
Sinningia Nees (>60), throughout the Neotropics
Vanhouttea Lem. (6), SE Brazil
Alloplectus Mart. (>75), throughout the Neotropics
(except SE Brazil)
Chrysothemis Decne. (7), Ecuador to Guatemala, C
Brazil, the Guianas, Venezuela and Lesser Antilles
Cobananthus Wiehler (1), Guatemala
Codonanthe (Mart.)Hanst. (>20), troughout the
Neotropics
Codonanthopsis Mansf. (5), Peru to Brazil (Aamazon
system)
Columnea L. s. lat.
Bucinellina Wiehler (2) , SW Colombia, NW Ecuador
Columnea L. s. str. (>75), C America, Carrib.
Islands, Colombia, Ecuador, Venezuala, Guayana Highlands, NE Brazil
Dalbergaria Tussac (>120), Bolivia to Guiana
Highlands, Mexico, Greater Antilles
Pentadenia (Planch.)Hanst. (>35), Bolivia to Mexico
Trichantha Hook.f. (>70), Bolivia to Guatemala,
Greater Antilles, Guayana Highlands
Corytoplectus Oerst. (15), Bolivia to Panama, coastal
Venezuela, Guayana Highlands
Drymonia Mart. (>140), throughout the Neotropics
Episcia Mart. s.l.
Episcia Mart. s str. (9), trop. S America to Nicaragua)
Alsobia Hanst. (2), Mexico and central America
Nautilocalyx [Linden ex] Hanst. (>70), troughout
trop. America, except SE Brazil
Nematanthus Schrad. (30), S & SE Brazil
Neomortonia Wiehler (3), central America and W Colombia
Oerstedina Wiehler (3), Central America
Paradrymonia Hanst. (>70), througout the Neotropcis,
except SE Brazil
Rhoogeton Leeuwenb. (3), British Guiana
Rufodorsia Wiehler (4), Costa Rica, Panama
Anetanthus Benth. (2), Peru, Bolivia, C Colombia, SE
Brazil
Besleria L. (>200), troughout the Neotropics, esp.
Andes of Colombia and Ecuador
Cremosperma Benth. (23), Costa Rica to Andes of Peru
Gasteranthus Benth. (35), S Mexico, central America, SW
South America to Bolivia
Reldia Wiehler (5), Panama to N Peru
Resia H.E.Moore (2), Colombia, Venezuela
Tylopsacas Leeuwenb. (1), Guayana Highlands
Napeanthus Gardn. (>30), troughout the Neotropics
Gesneria
L. s. lat.
Gesneria L. s. str. (Skog) (46), Carribean islands
Pheidonocarpa L. E. Skog (3), Carribean islands
Rhytidophyllum Mart. (c. 20), Carribean islands
Asteranthera Hanst. (1), Chile
Coronanthera C.B.Clarke (11), New Caledonia, one sp. in
Solomon Islands
Depanthus S.Moore (2), New Caledonia
Fieldia Cunn. (1), SE Australia
Lenbrassia G.W.Gillett (1), NE Australia
Mitraria Cav. (1), Chile, nearby Argentina
Negria F. Muell. (1), Lord Howe Island (SW Pacific)
Rhabdothamnus Cunn. (1), New Zealand
Sarmienta Ruiz & Pav. (1), Chile
Epithemateae
(= Klugieae) Klugieae Fritsch in Engl.
& Prantl, Nat. Pflanzenfam. 4(3b): 143. Mai 1893
Cyrtandromoea (11) SE Asia, Mal
Epithema Blume (> 20), S & SE Asia, one sp. in W
Africa
Gyrogyne W. T. Wang (1), S China
Loxonia Jack (3), W Malesia (Sumatra, Malay Penins.,
Borneo)
Monophyllaea R. Br. (32), Malesia (Sumatra to New
Guinea, S Thailand and Luzon to Java)
Rhynchoglossum Blume (c. 12), Malesia (India and S
China to New Guinea), 1 (or more?) spp. in C America, Columbia, Venezuela
Stauranthera Benth. (5-8), NE India and S China over
Malesia to New Guinea
Whytockia W. W. Sm. (6), S China, Taiwan
Acanthonema Hook.f. (2), trop. W Africa
Allocheilos W.T.Wang (2), S China
Allostigma W.T.Wang (1), S China
Ancylostemon Craib (11), C & S China
Anna Pellegr. (3), S China
Beccarinda Kuntze (c. 8), NE India, Burma, S China, Hainan, Vietnam, Sumatra
Boea [Comm. ex] Lam.(17), S China, NE India, N Thailand, Vietnam, Philipp. Sulawesi, Sumbawa, Flores, Ne Guinea, Solomon Islands, Bismarck and Lousiade Archipelago
Boeica C.B.Clarke (9), NE India, Bangla-desh, Burma, S
China, Malaysia
Bournea Oliv. (2), SE China
Briggsia Craib (21), C & S China
Briggsiopsis K.Y.Pan (1), S China
Calcareoboea H.W.Wu (1), S China
Cathayanthe Chun (1), S China
Championia Gard. (1), Sri Lanka
Chirita D.Don (>130), S China to Borneo
Chiritopsis W.T.Wang (8), S China
Colpogyne B.L.Burtt (1), Madagascar
Conandron Sieb. & Zucc. (1), China, Taiwan, Japan
Corallodiscus Batalin (17), India, Bhutan, Nepal,
Sikkim, C & S China
Dayaoshania W.T.Wang (1), S China
Deinocheilos W.T.Wang (2), S China
Deinostigma [W.T.Wang ex] Z.Y.Li (8), W Malesia
Didymocarpus Wall. (c. 80), S China to Malay Peninsula
and Sumatra
Didymostigma W.T.Wang (1), SE China
Dolicholoma D.Fang & W.T.Wang (1), SE China
Gyrocheilos W.T.Wang (4), S China
Haberlea Friv. (2), Balkan Peninsula
Hemiboea C.B.Clarke (21), C & S China, N Vietnam,
Taiwan, Japan
Hemiboeopsis W.T.Wang (1), S China
Henckelia Spreng. (Didymocarpus p.p., Loxocarpus,
Platyadenia) (c. 180), from S India and Sri Lanka troughout Malesia to New
Guinea
Hexatheca C.B.Clarke (3), Borneo
Hovanella A.Weber & B.L.Burtt (Didymocarpus p.p.)
(2), E & C Madagascar
Isometrum Craib (13), C China
Jancaea Boiss. (1), Greece
Lagarosolen W.T.Wang (2), S China
Leptoboea Benth. (4), NE India, Sikkim, Bhutan, Burma,
S China
Linnaeopsis Engl. (3), E Africa
Metabriggsia W.T.Wang (2), S China
Metapetrocosmea W.T.Wang (1), S China
Nodonema B.L.Burtt (1), trop. W Africa
Opithandra B.L.Burtt (9), S China, Japan
Orchadocarpa Ridl. (1), Malay Peninsula
Oreocharis Benth. (27), C & S China, N Vietnam,
Thailand
Ornithoboea [Parish ex] C.B.Clarke (11), S China to N
Malay Peninsula
Paraboea (C.B.Clarke) Ridl. (>80), Malesia (Sumatra
to New Guinea, S Thailand to Borneo)
Paraisometrum W.T.Wang (1), S China
Petrocodon Hance (1), China
Petrocosmea Oliv. (27), S China, NE India, Burma,
Thailand Vietnam
Phylloboea Benth. (1), Burma
Platystemma Wall. (1), Nepal, SW China
Primulina Hance (1), S China
Pseudochirita W.T.Wang (1), S China
Ramonda Rich. (3), Iberian (Pyrenees) and Balkan
Peninsula
Raphiocarpus Chun (Didissandra p.p.) (11), China,
Vietnam
Rhabdothamnopsis Hemsl. (1), S China
Ridleyandra A.Weber & B.L.Burtt (Didissandra p.p.)
(c. 23), Malay Peninsula, Borneo
Saintpaulia H.Wendl. (>20), E Africa
Schizoboea (Fritsch) B.L.Burtt (1), W, C & E Africa
Streptocarpus Lindl. (>130), Africa, Madagascar,
Comoro Islands
Tengia Chun (1), S China
Tetraphyllum [Griff. ex] C.B.Clarke (2-3), NE India,
Bangladesh, Thailand, ?Burma
Thamnocharis W.T.Wang (1), S China
Trachystigma C.B.Clarke (1), W Africa
Tremacron Craib (7), S China
Trisepalum C.B.Clarke (15), S China, Burma, Thailand, N
Malay Peninsula
Aeschynanthus Jack (c. 150), S China, Nepal, S India,
Sri Lanka, throughout Malesia to New Guinea, Solomon Islands
Agalmyla Blume (>50), Malesia
Loxostigma C.B.Clarke (7 or more), NE India, Sikkim,
Bhutan, Nepal, Burma, S China, N Vietnam
Lysionotus G.Don (c. 30) from N India and Nepal to Thailand,
Vietnam, S China, S Japan
Micraeschynanthus Ridl. (1, doubtful), Malay Peninsula
Cyrtandra J.R. & G.Forst. (>450), Nicobar
Islands, Sumatra, S Thailand, Taiwan, troughout Malesia to the Pacific and
Hawaiian Islands
Rhynchotechum Blume (12), SE Asia, position somewhat
uncertain, eventually to Didymocarpeae
Sepikaea Schltr. (1), New Guinea, probably to be
included in Cyrtandra
Genera of uncertain
position:
Sanango Bunting & Duke (1), C America, eastern
slopes of the Andes (Ecuador and Peru), included in Gesnerioideae-Gesnerieae by
Wiehler (1994), but position still doubtful
Titanotrichum Soler. (1), S China, Taiwan, S Japan
(accomodated in a monotypic tribe, Cyrtandroideae-Tiitanotricheae, by W.T.Wang)
Lamiaceae (= Labiatae) ( ~252/6700) Genera by subfamily and
tribe (after Cantino et al. 1992, 1997, Wagstaff et al. 1997, Olmstead, unpubl.
; Cantino pers. comm.).
Nomenclature follows Cantino et al 1997; 1998.
Teucrioideae (26/~1000)
Aegiphila Jacq. (125-150) trop Am
Ajuga L.
(40-50) mainly Eurasia
Amasonia L.f. (8) trop Am
Amethystea L. (1) Asia
Cardioteucris (= Rubiteucris) Kudo (2) China
Caryopteris Bunge (7) E Asia
Clerodendrum L. 400-500 trop/subtrop
Cyclonema
(= Rotheca)
Discretitheca
1 Nepal
Eplingia
(= Trichostema) L. (17) Mexico
Faradaya F. Muell. (3-18) Indomal/Austral/Polynes
Glossocarya Wall. ex Griff. (9-13) Australia/Indomal/SE
Asia
Hosea Ridl. (1) Indomal
Huxleya Ewart (1) Australia
Isanthus
(= Trichostema) N Am
Kalaharia
(= Clerodendrum) Africa
Karomia Dop. (9) Afr/Mad/Viet Nam
Kinostemon
(= Teucrium) China
Monochilus Fisch. & C.A. Mey. (2) Brazil
Oncinocalyx F. Muell. (1) Australia
Oxera Labill. (20) New Caledonia, Vanuatu
Pseudocaryopteris
3 S China, India, SE Asia
Rotheca
50-60 Afr/Mad/Ind Ocean/S Asia/Indomal
Rubiteucris
2 E Asia
Schnabelia Hand.-Mazz. (5) China
Spartothamnella Briq. (3) Australia
Tetraclea
(2) S. N.Am
Teucridium Hook.f. (1) New Zealand
Teucrium L. (200) worldwide mostly temp
Trichostema L. (17-19) N Am/Bahamas
Tripora 1
E Asia
Lamiina ()
Scutellarioideae Prantl, Text-Book Bot.: 293. 1 Mar-15 Apr 1880.
(5/372)
Harlanlewisia
(= Scutellaria) Ecuador
Holmskioldia Retz. (1) Himal
Perilomia
(= Scutellaria) S Am
Renschia Vatke (1) Africa
Salazaria
(= Scutellaria) SW US
Scutellaria L. (350) worldwide
Tinnea Kptschy & Peyr. (19) Africa
Wenchengia C.Y. Wu & S. Chow (1) China
Clebrookeina
Marrubiina
(~56/1090)
Achyrospermum Blume (10) Old World trop
Acrotome Benth. ex Endl. (8) trop and S. Africa
Ajugoides Makino (1) Japan
Alajja Ikonn. (3) C Asia/Afghan/W Himalaya
Ballota L. (35) Europe/Medit/W.Asia
Bostrychanthera Benth. (1) China
Brazoria Engelm. & A. Gray (2) S US
Chaiturus
(~ Leonurus) Ehrh. ex Willd. (1) W Eur to C As
Chamaesphacos Schrenk ex Firsh. & C.A. Mey. (1)
Iran/Afghanistan
Chelonopsis Miq. (13) Kashmir/Tibet/E Asia
Colquhounia Wall. (3) Himalaya/SW China
Craniotome Rchb. (1) Himalaya
Eremostachys Bunge (5+) Eur - C.Asia
Eriophyton Benth. (1) Himalaya
Galeobdolon
(= Lamium) W Eur/Iran
Galeopsis L. (10) temp Eurasia
Gomphostemma L. (30) SE Asia/Malaysia
Haplostachys (A. Gray) W.F. Hillebr. (6) Hawaii
Harmsiella
(= Otostegia)
Hypogomphia
4 C Asia/Afghan
Isoleucas O. Schwartz (1) Arabia
Lagochilus Bunge (35) C.As - Iran/Afghan.
Lagopsis Bunge (4) (=Marrubium?) Siberia/Japan
Lamiophlomis
(=Phlomis)
Lamium L. (41) N.Af/Eurasian
Lasiocorys
(= Leucas) trop S Af/Socotra/Arab
Leonotis (Pers.) R. Br. (40) trop, S Af; 1 pantrop
Leonurus L. (4) temp Eurasia
Leucas R. Br. (150) Afr/Arabia/Indomal.
Loxocalyx Hemsl. (2) China
Macbridea Elliott ex Nutt. (2) SE US
Marrubium L. (30) Eur/Medit
Melittis L. (1) Eur - Iran
Metastachydium Airy Shaw ex C.Y. Wu & H.W. Li (1)
C. Asia
Microtoena Prain (25) Himalaya-W China
Moluccella L. (4) Medit - NW India
Notochaete Bent. (1) Himalaya
Otostegia Benth. (20) SE trop Af - C Asia
Panzerina Sojak (6) Siberia/Mongol
Paralamium Dunn (1) SW China
Paraphlomis Prain (8) E Asia/Malaysia
Phlomidoschema (Benth.) Vved. (1) Iran-W Himalaya
Phlomis L. (100) C Asia/China
Phyllostegia Benth. (25) Tahiti/Hawaii
Physoleucas
(= Leucas) Arabia
Physostegia Benth. (12) N Am
Prasium L. (1) Medit
Pseuderemostachys Popov (1) C Asia
Pseudomarrubium Popov (1) C Asia ?
Roylea Wall. ex Benth. (1) Himalaya
Sideritis L. (100) N temp Old world/Macarones
Stachyopsis Popov & Vved. (3) C Asia
Stachys L. (300) temp World-wide
Stenogyne Benth. (25) Hawaii
Sulaimania Hedge & Rech.f. (1) Pakistan
Suzukia Kudo (2) Ryukyus/Taiwan
Synandra Nutt. (1) E US
Thuspeinanta T. Durand (2) C Asia/Iran/Afghan
Warnockia
1 S US
Wiedemannia Fisch. & C.A. Mey. (3) SW Asia
Colebrookeina genera of
uncertain placement
Anisomeles R. Br. (13) OW trop, Mad, Indom, Aus
Colebrookea Sm. (1) India
Comanthosphace S. Moore (5) E Asia
Epimeredi
(= Anisomeles)
Eurysolen (or Lamioideae?) Prain (1) SE Asia and
Malaysia
Eusteralis
(= Pogostemon) 96 temp Asia/Aus
Leucosceptrum Sm. (3) Himal., China
Pogostemon Desf. (71) Indomalaysia
Rostrinucula Kudo (2) E Asia
Holocheila (Kudo) S. Chow (1) China (= Teucrieae?)
Prostantheroideae (16/239) Leurss., Handb. Syst. Bot. 2: 1014, 1016. Sep 1882 (14/238)
Chloantheae (9/99) Benth. & Hook.f., Gen. Pl. 2: 1132. 1-16 Mai 1876
Chloanthes R. Br. (4) Australia
Cyanostegia Turcz. (5) Australia
Denisonia (=Pityrodia)
Dicrastylis J.L. Drumm. ex Harv. (26) Australia
Hemiphora (F. Muell.) F. Muell. (1) Australia
Lachnostachys Hook. (6) Australia
Mallophora Endl. (2) Australia
Newcastelia F. Muell. (12) Australia
Physopsis Turcz. (2) Australia
Pityrodia R. Br. (41) Australia
Westringieae
Bartl., Ord. Nat. Pl.: 182. Sep 1830. (7/140)
Cryphia
(= Prostanthera)
Eichlerago
1 (= Prostanthera) Australia
Hemiandra R. Br. (7) Australia
Hemigenia R. Br. (40) Australia
Microcorys R. Br. (16) Australia
Prostanthera Labill. (45) Australia
Westringia Sm. (30) Australia
Wrixonia 1
Australia
Symphorematina (3/24)
Congea Roxb. (7) SE Asia & W. Malaysia
Sphenodesme Jack (14) SE Asia & W. Malaysia
Symphorema Roxb. (3) Indomal
Nepetoideae
Kostel., Allg. Med.-Pharm. Fl. 3: 752. Apr-Dec 1834. (/)
Mentheae
Dumort., Fl. Belg.: 48. 1827. (/)
Acanthomintha A. Gray (3) W. US
Acinos (= Clinopodium)
Agastache Gronov. (22) N. Amer.
Blephilia Raph. (3) E N Am
Bystropogon L'Her. (10) Macaronesia
Calamintha (= Clinopodium)
Cedronella Moench (1) Macaronesia
Chaunostoma Donn.Sm. (1) C. Amer.
Cleonia L. (2) W Medit.
Clinopodium L. (100) N World temp Eurasia
Conradina (6) SE US
Coridothymus (= Thymbra) Rchb.f. (unknown) Uncertain
Cuminia Colla (1) Juan Fernandez Isl.
Cunila Royen ex L. (15) N/S Am
Cyclotrichium (Boiss.) Manden. & Scheng. (6) SW
Asia to Iran
Dicerandra Benth. (8-9) SE US
Diodeilis (= Clinopodium)
Dorystaechas Boiss. & Heldr. ex Benth. (1) W Asia
Dracocephalum L. (46) Eurasia, N. Amer.
Drepanocaryum Pojark. (1) C Asia
Eriothymus (Benth.) Rchb. (1) Brazil
Euhesperida Brullo & Furnari (1) Libya &
Cyrenaica
Gardoquia (= Clinopodium)
Glechoma L. (10) temp. Eurasia
Glechon Spreng. (6) Brazil, Paraguay
Gontscharovia Boriss. (1) C Asia
Hedeoma Pers. (38) SW N. Amer. & S. Amer.
Hesperozygis Epling (8) Mex. to Brazil
Hoehnea Epling (4) S Brazil to N Argentina
Horminum L. (1) S Europe
Hymenocrater Fisch. & C.A. Mey. (12) SW Asia
Hyssopus L. (5) S Europe & Medit. to C Asia
Kudrjaschevia Pojark. (4) C Asia
Kurzamra Kuntze (1) temp. S. Amer.
Lallemantia Fisch. & C.A. Mey. (5) Turkey to C Asia
& Himal.
Lepechinia Willd. (55) warm N. Amer. & S. Amer.,
Hawaii
Lophanthus Adans. (2) unknown
Lycopus L. (14) N temp
Marmoritis Benth. (5) Pakistan to China
Meehania Britton (2) E Asia, E US
Melissa L. (3) Europe to Iran & C Asia
Mentha L. (25) temp. OW
Meriandra Benth. (2) Ethiopia, Himal.
Micromeria Benth. (90) temp. Europe, Canary Isl., W.I.
Minthostachys (Benth.) Spach (12) W S. Amer.
Monarda L. (20) N Am, Mex
Monardella Benth. (20) W N. Amer.
Neoeplingia Ramam., Hiriart & Medrano (1) Mex.
Nepeta L. (250) Eurasia & N Africa
Obtegomeria (1) N S Am
Origanum L. (36) Eurasia
Pentapleura Hand.-Mazz. (1) Kurdistan
Perovskia Kar. (7) NE Iran to NW India
Piloblephis Raf. (1) SE US
Pitardia Batt. ex Pit. (2) NW Africa
Pogogyne Benth. (6) W N. Amer.
Poliomintha A. Gray (7) SW N. Amer.
Prunella L. (4) N temp. Eurasia & NW Africa
Pycnanthemum Michx. (17-21) E N Am
Rhabdocaulon (Benth.) Epling (7) trop. S. Amer.
Rhododon Epling (1) SC US
Rosmarinus L. (2) Medit.
Saccocalyx Coss. & Durieu (1) NW Africa
Salvia L. (900) OW & NW temp regions
Satureja L. (30) OW & NW temp. & warm regions
Schizonepeta (Benth.) Briq. (3) temp. Asia
Stachydeoma Small (1) SE US
Thymbra L. (4) Medit. & S Europe to SW Asia
Thymus L. (350) temp. Eurasia
Xenopoma (= Clinopodium)
Zataria Boiss. (1) Iran, Afghanistan
Zhumeria Rech.f. & Wendelbo (1) Iran
Ziziphora L. (25) Medit. to C Asia
Ocimeae Dumort., Anal. Fam. Pl.: 22. 1829 (52/1458)
Acrocephalus Benth. (= Platostoma)
Aeollanthus Mart. ex Spreng. (43) trop. & warm
Amer.
Alvesia Welw. (3) trop. Africa
Anisochilus ()
Asterohyptis Epling (3) Mex.
Basilicum Moench (7) OW trop. to Australia
Becium (= Ocimum) Lindl. (10) trop. Africa
Benguellia G. Taylor (1) Angola
Capitanopsis S. Moore (2) Madag.
Capitanya Schweinf. ex Gurke (1) E Africa
Catoferia (Benth.) Benth. (4) C. Amer.
Ceratanthus (= Platostoma) F. Muell. ex G. Taylor (10)
SE Asia, Malayasia
Dauphinea Hedge (1) Madag.
Endostemon N.E. Br. (17) trop. & S Africa to India
Englerastrum (= Plectranthus) Briq. (20) trop. Africa
Eriope Kunth ex Benth. (30) trop. & warm S. Amer.
Eriopidion (= Eriope) Harley (1) Venezula, Brazil
Erythrochlamys (= Ocimum) Gurke (5) trop. E Africa
Fuerstia T.C.E. Fr. (6) trop. Africa
Geniosporum (= Plectranthus) Wall. ex Benth. (25) trop.
Africa, Madag., SE Asia
Hanceola Kudo (3) China
Haumaniastrum P.A.Duvign. & Plancke (23) trop.
Africa
Hemizygia (Benth.) Briq. (28) trop. & S Africa
Holostylon Robyns & Lebrun (3) trop. Africa
Hoslundia Vahl (2) trop. Africa
Hypenia (Mart. ex Benth.) Harley (24) warm & trop.
Amer.
Hyptidendron Harley (16) warm & trop. Amer.
Hyptis Jacq. (300) warm & trop. Amer., OW trop.
Isodictyophorus (= Plectranthus) Briq. (1) W Africa
Isodon (Schrad. ex Benth.) Spach (96) warm & trop.
Asia & Africa
Leocus ()
Marsypianthes Mart. ex Benth. (6) trop. Amer.
Mesona (= Plectranthus) Blume (3) Asia, Malaysia
Neohyptis (= Plectranthus) J.K. Morton (1) Angola
Nosema (= Plectranthus) Prain (6) SE Asia, Malaysia
Ocimum L. (150) temp. & trop. OW
Octomeron (= Plectranthus) Robyns (1) trop. Africa
Orthosiphon Benth. (40) OW trop.
Peltodon Pohl (5) Brazil, Paraguay
Perrierastrum Guillaumin (1) Madag.
Platostoma P. Beauv. (5) OW trop.
Plectranthus L'Her. (200) warm & trop. OW
Puntia Hedge (1) C Somalia
Pycnostachys Hook. (37) trop & S Africa, Madag.
Rabdosiella (= Plectranthus) Codd (2) S Africa, N India
Rhaphiodon Schauer (1) Brazil
Solenostemon (= Plectranthus) Thonn. (10) trop. Africa
Symphostemon (= Plectranthus) Hiern (2) Angola
Syncolostemon E. Mey. (10) SE Africa
Tetradenia Benth. (9) trop. Africa, Madag.
Thorncroftia N.E. Br. (3) Transvaal
Elsholtzieae
Burnett, Outl. Bot.: 971. Jun 1835 (6/57)
Collinsonia L. (4) E N Am
Elsholtzia Willd.(35) OW trop.
Keiskea Miq. (5) Asia
Mosla (Benth.) Buch.-Ham. ex Maxim. (10) E Asia,
Malaysia
Perilla L. (6) India to Japan
Perillula Maxim. (1) Japan
Lavanduleae Caruel in Parl., Fl. Ital. 6: 53. Sep 1884
(1/30)
Lavandula L. (30) Atlantic isl., Medit. to Somalia
& India
Sabaudia (= Lavandula) Buscal. & Muschl. (2) NE
& SE trop. Africa
Nepetoideae genera of
uncertain placement
Bovonia Chiov. (1) Zaire
Heterolamium C.Y. Wu (1) China
Ombrocharis Hand.-Mazz. (1) China
Viticoideae (3/260)
Petitia Jacq. (2) W.I.
Pseudocarpidium Millsp. (8) W.I.
Teijsmanniodendron Koord. (14) SE Asia, Malaysia
Vitex L. (250) temp. & trop. NW & OW
Acrymia Prain 1 Malaysia
Adelosa Blume
1 Madagascar
Archboldia
E. Beer & H.J. Lam (1) New Guinea
Callicarpa
L. (140) warm & trop. NW & OW
Cornutia
L. (12) trop. Am
Cymaria
Benth. (2-3) SE Asia, Indomal.
Garrettia
H.R. Fletcher (1) SE Asia, Indomal.
Geunsia (=Callicarpa)
Gmelina L.
(35) SE Asia, Indomal., Africa, Mascarenes
Hymenopyramis Wall. ex Griff. (6) India, SE Asia
Neorapinia Moldenke (1) New Caledonia
Paravitex H.R. Fletcher (1) SE Asia
Peronema Jack (1) SE Asia/Indomal.
Petraeovitex Olive (8) Indomal /Pac.Is/Phillipines
Premna 200 trop/subtrop Af/Asia/Aust
Pseudochamaesphacos ?
Pygmaeopremna (= Premna?)
Tectona L.f. (3) trop. Asia
Tsoongia Merr. (1) SE Asia
Viticipremna Lam. (5) Indomal., Australia
Lentibulariaceae Rich. in Poit. & Turpin, Fl. Paris 1: ed. fol.
23, ed. qu. 26. 1808, nom. cons.
(4/310) See Mller et al.
2004.
Byblis (2) Aus, New Guinea
Genlisea A. St.-Hil. (21) trop Am, trop & S Af, Mad
Pinguicula L. (74) Am, Medit, circumbor
Utricularia L. (215) cosmop, esp trop
Linderniaceae (Rchb.) Borsch, K. Mller, & Eb. Fisch. (13/~194)
Artanema D. Don (4) trop Afr, Asia
Chamaegigas Dinter (1) Namibia
Craterostigma Hochst. (9) Yemen, trop & S Af, Ind
Crepidorhopalon Eb. Fisch. (28) trop & S Af, Mad
Hartliella Eb. Fisch. (4) C Af (Congo, Katanga)
Hemiarrhena Benth. (1) Qland (= Lindernia?)
Legazpia Blanco (1) SE As, Micrones, N Guin (=
Torenia?)
Lindernia All. (~100) Eur, trop Af, Mad, SE As, N, C,
& C. Am.
Picria Lour. (1) Indomal. (= Curanga)
Pierranthus Bonati (1) SE As
Schizotorenia T. Yamaz. (2) SE As (= Lindernia?)
Scolophyllum T. Yamaz. (2) SE As (= Lindernia?)
Torenia L. (40) SE As, Af, Mad
Martyniaceae Horan., Char. Ess. Fam.: 130. 1847, nom. cons. (4/17) (sensu H.-D. Ihlenfeldt, pers. comm.)
Craniolaria
L. (3) S. Am.
Ibicella
Van Eselt. (3) warm and trop. Am
Martynia
L. (1) Mex
Proboscidea
Schmidel (10) warm Am
Oleaceae Hofmanns. & Link Fl. Portug. 1:62. 1809. (25/600+) provided by Eva Wallander
(eva.wallander@systbot.gu.se) 6/14/00
Comments: Note that Dimetra and Nyctanthes now, without doubt, belong to Oleaceae (not Verbenaceae). The
subfamilies Jasminoideae and Oleoideae have been abandoned because Jasminoideae
is paraphyletic. The Oleoideae is monophyletic but now recognised as tribe
Oleeae, with its former tribes as subtribes.
Fontanesieae
Fontanesia (1-2) SW Asia (and Sicily) and China
Forsythieae
Abeliophyllum
Nakai (1) Korea
Forsythia
(11) E Asia and SE Europe (1 sp.)
Jasmineae
Jasminum (200+) Tropical and subtropical parts of the
Old World
Menodora (22) Subtropical North America, central and S
South America, and S Africa
Myxopyreae
Dimetra
(1) Thailand
Myxopyrum (4) Tropical SE Asia
Nyctanthes (2) Tropical and subtropical SE Asia
Oleeae
Ligustrinae
Ligustrum (45) Temperate to tropical parts of the Old
World, except Africa
Syringa (20) Mainly subtropical parts of Eurasia
Schreberinae
Comoranthus
(3) Madagascar and the Comores
Schrebera
(4) Tropical parts of Africa and India
Fraxininae
Fraxinus (50) Mainly temperate and subtropical regions
of the Northern Hemisphere
Oleinae
Chionanthus (100) Tropical and subtropical Africa,
America, Asia, and Australia
Forestiera (15) Subtropical North America, West Indies,
and N South America
Haenianthus (3) West Indies
Hesperelaea (1, extinct) Mexico
Nestegis (5) New Zealand and Hawaii (1 sp.)
Noronhia (41) Madagascar
Notelaea (12) Australia and Tasmania
Olea (40+) Tropical and subtropical parts of the Old
World
Osmanthus (30) Subtropical parts of E Asia and North
America (1-2 spp.)
Phillyrea (2) Mediterranean region to W Asia
Picconia (1) Macaronesia
Priogymnanthus
(2) South America
Orobanchaceae Vent., Tabl. Rgne Vg. 2:292. 1799, nom. cons. (65/1538) Synoptical classification following Young
et al. 1999, Olmstead et al. 2001, Wolfe et al. 2005, Bennett & Mathews,
unpubl., and D. Nickrent, pers. comm.
Orobanchaceae sensu stricto
Boschniakia (4) Himal, N Asia, NW N Am
Cistanche (16) Med, Ethiopia, India, to W China
Conopholis (2) E US to Panama
Diphelypaea (2?) Asia
Epifagus (1) E N Am
Kopsiopsis (2) W N Am (= Boschniakia)
Mysorrhiza (= Orobanche)
Orobanche (100-150) temp & warm N Hemisphere
Xylanche
(2) Himal to Taiwan (= Boschniakia)
New World hemiparasite plus Pedicularis clade
Agalinis (40) trop & warm Am
Adenostegia (= Cordylanthus)
Castilleja (200) most W N Am, also E N Am, Euras, C Am,
Andes
Clevelandia (1) Baja Ca (= Castilleja)
Cordylanthus (18) W N Am
Esterhazya (5) Bolivia, Brazil
Gentrya (= Castilleja)
Gerardia (= Agalinis)
Lamourouxia (28) Mex to Peru
Macranthera (1) SE US
Ophiocephalus (1) Baja Ca (= Castilleja)
Orthocarpus (9) W Am
Pedicularis (350) N hemisphere, 1 sp in Andes
Phtheirospermum (7) E As
Seymeria (25) S N Am
Triphysaria (5) W N Am
Africa/SW Asia clade
Aeginetia L. (3) Indomal, E As
Alectra (40) Africa/Asia
Buchnera (100) warm O&NW (16 NW)
Cycnium (15) warm AF (~Escobediea)
Escobedia (6) trop Am
Graderia (5) Af & Socotra
Harveya (40) trop & S. Af, Mascarenes
Hyobanche (7) S Af
Melasma (20) trop Af, Am
Radamaea (5) Madagascar
Sopubia (60) trop OW to S Af
Stellularia (= Buchnera)
Striga (~40) OW trop to S Af
Vellosiella (3) Brazil (~Melasma)
Xylocalyx (5) Somalia, Socotra
Cyclocheilaceae
Asepalum (1) NE & E Af
Cyclocheilon (3) NE Af
Nesogenaceae
Nesogenes A. DC. (8) E Afr, Mad, India, Pac Is
European clade
Bartsia (60) Eur, Af mt, Andes (not mono-
Bennett&Mathews)
Bellardia (1) Med
Bungea (2) SW As to China
Euphrasia (~170) N temp, Malaysia, NZ, S Am (only
Eurasia? Nickrent)
Fistularia (= Rhinanthus)
Lathraea (7) Eurasia
Macrosyringion (= Odontites)
Melampyrum (35) N temp
Odontites (30) W & S Eur to W As
Orthantha (= Odontites)
Parentucellia (2) Med & SW As
Rhynchocorys (6) S Eur to Iran
Rhinanthus (45) N hemisphere, esp Eur
Siphonidium (= Euphrasia)
Tozzia (1) Alps, Carpathanians
Orobanchaceae placed in tree, but unassigned to above
clades
Brandisia (13) Burma China
Cymbaria (4) Eurasia
Lesquereuxia (1) E Medit (= Siphonostegia?)
Lindenbergia (15) Himal to trop Afr/Asia
Monochasma (4) E As
Schwalbea (1) E US
Siphonostegia (3) Med & E As
Genera not sampled in phylogenetic studies and
unassigned for that reason.
Buchnereae (25/352)
Aureolaria (11) E US, Mex
Buttonia (3) trop & S Af
Centranthera (9) China
to Australia
Gerardiina (2) trop & S. Af
Leptorhabdos (1) Cauc, Iran, C As, Himal
Micrargeria (4) trop Af, India
Nothochilus (1) Brazil
Physocalyx (2) Brazil
Rhamphicarpa (6) trop & S Af, India, Austral
Rhaphispermum (1) Madag (=Pseudomelasma?)
Silviella (2) Mex
Tetraspidium (1) Madagascar
Orobancheae (15/210)
Christisonia (17) SW China, SE As
Gleadovia (6) W Himal, W. China
Mannagettaea (3) W China to E Siberia
Necranthus (1) Turkey
Phacellanthus (1) E As
Phelypaea (4) Medit
Platypholis (1) Japan
Tienmuia (1) E China (~Phacellanthus)
Rhinantheae (21/944)
Omphalotrix (1) NE As
Unassigned:
Ancistrostylis (1) Laos
Brachystigma (1) SW US
Cycniopsis (3) trop Africa
Dasistoma (1) E N Am
Ghikaea (1) trop Africa
Hedbergia (1) trop Africa
Leucosalpa (3) Madagascar
Madgalenaea (1) SE Brazil
Micrargeriella (1) trop E Africa
Nothobartsia (1) Medit
Parastriga (1) trop Africa
Petitmenginia (2) S China, SE Asia
Pseudobartsia (1) China
Pseudomelasma (1) Madagascar
Pseudosopubia (7) trop Africa
Pseudostriga (1) SE Asia
Pterygiella (3) China
Sieversandreas (1) Madagascar
Spirostriga (1) California to Baja Ca
Thunbergianthus (2) Africa
Vellosiella (3) Brazil
Zizangia (1) China
Paulowniaceae Nakai J. Jap. Bot. 24:13, 1949. (1/6)
Paulownia
Siebold & Zucc. (6) E Asia
Pedaliaceae R.Br. Prodr. 519. 1819. nom.cons.
(13/70) (sensu H.-D. Ihlenfeldt, pers. comm.. 2001)
Pedalioideae (of Pedaliaceae s.l.) 12/70
Ceratotheca
(5) trop and S. Af.
Dicerocaryum
(3) S. trop. Af.
Harpagophytum
(2) S. Af. Madagascar
Holubia
(1) S. Af.
Josephinia
(4) Kenya to Somalia
Linariopsis
(2-3) W and SW trop Af.
Pedaliodiscus
(1) E Af
Pedalium
(1) OW trop P. murex edible
Pterodiscus
(13) trop and S. Af.
Rogeria
(4) Brazil, trop and S. Af.
Sesamothamnus
(6) trop Af.
Sesamum
(19) OW trop. And S. Af.
Uncarina
(13) Madagascar
Berendtiella
(4) C Am
Dodartia
(1) S Russia, W As
Elacholoma (1) C Aus
Glossostigma (6) Aus, NZ, Af, Ind
Hemichaena
(1) C Am
Lancea
(2) Tibet, China
Leucocarpus
(1) trop Am
Mazus
(20) E & SE As, Indomal, Aust
Mimulus
(150) Am, Asia, S Af
Peplidium (7) Aus, Ind, Af
Phryma
(2) E Asia, E N Am
Plocospermataceae Hutch. Fam. Fl. Pl. ed. 3 469. 1973. (1/1)
Plocospermum
(1) Mex, Guatemala
Schlegeliaceae (A.H. Gentry) Reveal Phytologia 79:74.
1996. (3/20) formerly of
Bignoniaceae and may still be close
Gibsoniothamnus
(7) Mexico to C Am
Schlegelia
(12) tropical Am
Synapsis
(1) Cuba
Thomandersia
Baill. (6) trop Af
Scrophulariaceae Durande
Notions Elm. Bot.: 265.
1782. (52/1681) Synoptical
classification by tribe. Following Wettstein (1891), Barringer (1993), Olmstead
et al. 2001, Kornhall et al., 2001, 2004, Oxelman et al. 2005.
Aptosimeae (4/44)
Anticharis
(14) Af to Mal
Aptosimum
(20) Africa
Peliostomum
(7) trop & S Af
Stemodiopsis
(3) Africa
Buddlejaceae (5/~110)
Buddleja (98) Neotrop (63) S Af (10) As (25)
Chilianthus (?) ?
Emorya
(2) Tex., Mex.
Gomphostigma
(2) S. Af.
Nicodemia
(=Buddleja?)
Freylinieae (4/8) (Barringer 1993)
Antherothamnus
(1) C & S Af
Freylinia
(4) trop & S Af
Manuleopsis
(1) SW Afr
Phygelius
(2) S Africa
Hemimerideae (7/134)
Alonsoa
Ruiz & Pav. (16) trop Am, S Af
Colpias
(1) S Africa
Diascia
(38) S Af
Diclis
(10) trop & S Af, Mad
Hemimeris
(4) S Af
Nemesia
(65) trop & S Af
Schistanthe
(?) ?
Leucophyleae (3/17)
Capraria
(4) warm Am
Eremogeton
(1) Mex, Guat
Leucophyllum
(12) SW N Am
Manuleeae (28/566) (Hilliard 1994)
Agathelpis
Choisy (2) SW Cape
Barthlottia (1) Madag
Chenopodiopsis (3) S Af
Cromidon
(12) S Af
Dischisma
(11) SW Cape to Namibia
Glekia (1) S Af
Globulariopsis
(7) S. Af
Glumicalyx (6) S&SE Af
Gosela
(1) S Af
Hebenstretia
(25) trop & S Am
Jamesbrittenia
(83) S Af
Limosella (11) cosmop
Lyperia
(6) S Af
Manulea
(73) S Af
Manuleopsis
(1) SW Af
Melanospermum
(6) S Af
Microdon
(7) SW Cape
Phyllopodium
(26) SW Cape
Polycarena
(17) SW Cape
Pseudoselago
(28) west S Af
Reyemia
(2) S Af
Selago
(190) trop & S Af
Sphenandra?
(= Sutera)
Strobilopsis
(1) S Af
Sutera
(49) S Af
Tetraselago
(4) S Af
Trieenia
(9) SW Cape
Walafrida
(?) Madag, S Af
Zaluzianskya
(55) W S Af
Myoporeae (4/236) sensu Chinnock, in press
Androya (1) Madagascar (Oxelman et al. 1999)
Bontia
L. (1) W. Ind., trop S. Am.
Calamphoreus
Chinnock (1)
Diocirea
Chinnock (4) Austral
Eremophila
R. Br. (214) Austral. (1 NZ)
Glycocystis
Chinnock (1) Austral
Myoporum Sol. Ex Forster (30) Austral., NZ, Pac. Is.,
E. Asia
Pentacoelium Zucc. (1) Austral
Scrophularieae (4/564)
Antherothamnus
(1) C&S Af
Celsia
(= Verbascum)
Oreosolen (3) Himalaya
Scrophularia
(200) N temp to trop Am
Staurophragma
(= Verbascum)
Verbascum
(360) Euras, E Af
Teedieae (3/6) (G. Don; Barringer 1993)
Oftia
(3) S Af
Ranopisoa
(1) Madag (~ Oftia)
Teedia
(2) S Af
Unassigned
Androya
(1) Mad
Camptoloma (3) Canaries Ethiopia Arabia
Cheilophyllum
(8) W Indies
Dermatobotrys
(1) S. Af
Stilbaceae (11/39) Synoptical Classification by tribe. (Bremer et al. 1994; Olmstead et al.
2001; Oxelman, unpubl.; Goldblatt, pers. comm.)
Stilbeae 7/28
Campylostachys
(1) SW Cape
Eurylobium
(2) SW Cape
Euthystachys
(1) S Af
Nuxia
(15) S Arabia to trop Af, Masc, S Af.
Retzia
(1) Cape
Stilbe
(6) SW and S Cape
Xeroplana
(2) SW and S Cape
Bowkerieae (Barringer 1993) 3/7
Anastrabe
(1) S Africa
Bowkeria
(5) S Africa
Ixianthes
(1) S Africa
Hallerieae 1/4
Halleria
(4) trop & S Africa, Madag
Polypremum
(1) warm Am.
Tetrachondra
(2) Patagonia, N.Z.
Casselieae (4/25)
Casselia
12 trop Am
Ghinia
7 trop Am
Parodianthus 2 Argentina
Tamonea
4-5 trop Am
Citharexyleae (6/108)
Baillonia 1 S.Am
Citharexylum 70 trop Am
Duranta 30 Carib./S.Am
Recordia 1 Bolivia
Rehdera 3 C.Am
Rhaphithamnus 2 Chile/Juan Fern.
Verbenoxylum
1 Brazil
Petreeae (2/12)
Petrea 11 (Rueda 94 only 11) trop Am
Xolocotzia 1 Mexico
Priveae (3/22)
Castelia
(=Pitraea)
Dipyrena 1 temp S Am
Pitraea
(~Priva) 1 S Am
Priva 20 trop/subtrop Af/Am
Lantaneae (12/497)
Acantholippia 6 arid S.Am
Aloysia
37 New World
Bouchea 9 trop Am
Burroughsia
(= Lippia) 2 SW N Am.
Chascanum
23 Af/Mad./Arab./Ind.
Diostea 3 S Am
Gonyostachyum
(~Lippia, Sanders 2001)
Lampaya 3 S. Am deserts
Lantana 150 trop Am/trop; S Af
Lippia 200 trop Af/Am
Nashia 7 Bahamas/Cuba
Neosparton 4 temp. S.Am
Riedela
(~Lippia, Sanders 2001)
Phyla
(~Lippia) 15 trop Am
Stachytarpheta
65 trop Am/trop/Af
Xeroaloysia
1 Argentina
Verbeneae (6/311)
Glandularia
~50 S Am/N Am, Mex
Hierobotana 1 Ecuador/Peru
Junellia 57 S. Am.
Stylodon 1 SE US (= Verbena)
Urbania 2 Chile
Verbena ~200 mostly New World
Svensonia
(~Chascanum) 4 E.Af/India
Ubochea
(~Stachytarpheta) 1 C Verde I
Unplaced to tribe (2/2-3)
Coelocarpum 1 Socotra/Madagascar
Saccanthus
(1-2) Bolivia
Veronicaceae (= Plantaginaceae in APGII) (104/1819) Synoptical classification by
Tribe. Following Wettstein (1891),
Barringer (1993), Olmstead et al. 2001, Oxelman et al. 1999, Oxelman et al.
2005; Albach et al. 2005.
Angelonieae (5/39)
Angelonia
(25) trop Am
Basistemon
(8) trop Am
Melosperma
(1) Chile
Monopera
(2) S. Am
Monttea
(3) Chile
Antirrhineae 27/326 (Sutton 1988)
Acanthorrhinum (1) N Af
Albraunia (3) SW Asia
Anarrhinum (8) Medit. SW Asia
Antirrhinum (20 Sutton - Medit; 36 Thompson Medit, W
N Am)
Asarina (1) SW Europe
Chaenorrhinum (21)
Medit.
Cymbalaria (9) W Europe to Iran
Epixiphium (1) SW N Am (~ Maurandya)
Galvezia (4) coast Peru, Ecuador, Galap.
Gambelia (4) SW N Am (~Antirrhinum)
Holmgrenanthe (1) SW Nam
Holzneria (2) SW Asia
Howelliella (1) SW N Am (~ Antirrhinum)
Kickxia (46) Med to SC&SW Asia
Linaria (150) Euras, Medit
Lophospermum (8) N&C Am
Mabrya (5) SW N Am
Maurandella (1) SW N Am (~Maurandya)
Maurandya (2) Mex C Am
Misopates (7) Medit, Cape Verde, Is Ethiop India
Mohavea (2) SW US
Neogaerrhinum (2) (~Antirrhinum)
Nuttalanthus (4) N Am temp S Am (~Linaria)
Pseudorontium (1) SW N Am (~Antirrhinum)
Rhodochiton (3) Mex (~Lophospermum)
Sairocarpus (13) SW US (~ Antirrhinum)
Schweinfurthia (6) NE Af to India
Callitricheae 2/18
Callitriche
(17) Cosmop
Hippuris
(1) Cosmop
Cheloneae 8/318
Brookea (3) Borneo
Chelone (4) E N Am
Chionophila (2) W N Am
Collinsia (20) N Am
Keckiella (7) N Am
Nothochelone (1) W N Am
Pennellianthus (1) NE As
Penstemon (275) N Am
Tonella (2) W US
Uroskinnera (3) Mex C Am
Digitalideae 3/24
Digitalis (19) Med, Eur, C As
Erinus (2) N Af, Pyrren, Alps
Isoplexis (3) Macaronesia
Globularieae 2/24
Globularia (22) Cape Verde Eur W As
Poskea (2) Somolia
Socotra
Gratiolieae 25/271
Achetaria Cham. & Schltdl. (5) trop Am
Adenosma R. Br. (15) China, Indomal, Aus
Allocalyx?
(=Monocardia?)
Amphianthus? (=Bacopa)
Artenema (4) OW trop
Bacopa (56) warm esp Am
Benjaminia (1) trop Am
Bryodes (3) Mascaren
Bythophyton? (1) Indomal
Conobea (7) Am
Dintera (1) trop Af
Dizygostemon (1) Brazil
Dopatrium (7) trop As, Aus
Encopella (1) Cuba
Geochorda (=Bacopa)
Gratiola (20) temp, trop mts
Hemianthus (=Micranthemum)
Hemisiphonia (=Micranthemum)
Herpestis (=Bacopa)
Hydrotriche? (4) Mad
Ildefonsia (=Bacopa)
Leucospora (=Stemodia?)
Limnophila (37) OW trop
Mecardonia (10) warm
Am (~Bacopa)
Micranthemum (14) Am, WI
Microcarpaea (=Peplideum)
Monocardia (=Bacopa?)
Morgania (= Stemodia)
Otacanthus (4) Brazil
Psammetes (1) trop W Af
Schistophragma? (2) Am (~Leucocarpus)
Scoparia (20) trop Am
Stemodia (55) OW & NW trop
Tetraulacium (1) Brazil
Trapella (1-2) E. As.
Zenkerina? ()
Hemiphragmieae 1/1
Hemiphragma (1) Himal
Plantagineae (3/285)
Aragoa (8) Andes
Bougueria
(1) Andes (= Plantago)
Littorella (3) 1 each in N Am, S Am, Eur
Plantago
(273) cosmop
Russelieae 2/54
Russelia
(52) Cuba, Mex to Columbia
Tetranema (2) Mex Guatem
Sibthorpieae 2/6
Ellisiophyllum
(1) India to Japan, New Guinea
Sibthorpia (5) trop & S Am, Macaron, Eur, Afr mts
Veroniceae 21/400 (see Albach et alo. 2004)
Besseya
(7) N Am (= Veronica)
Botryopleuron (=Veronicastrum)
Calorhabdos (=Veronicastrum)
Chionohebe
(6) Aus, NZ (= Veronica)
Cochlidiosperma (= Veronica)
Derwentia (10) SE Aus (= Veronica)
Detzneria (1) New Guin (= Veronica)
Falconeria
(=Kashmiria)
Hebe (95) NZ, Fr Polynes, S Am (= Veronica)
Heliohebe (5) NZ (= Veronica)
Kashmiria (1) Himal
Lagotis (20) Eur to W China
Neopicrorhiza (2) E Himal (= Picrorhiza)
Odicardis (2) Cauc, E Afr (= Veronica)
Oligospermum (= Veronica)
Paederota (2) S Eur
Paederotella (1-3) As Minor, Cauc (= Veronica)
Parahebe (30) N Guin,ea NZ (= Veronica)
Picrorhiza (3) W Himal
Pseudolysimachion (19) temp/subtemp Euras (= Veronica)
Scrofella (1) China
Synthyris (9) W N Am (= Veronica)
Veronica (s.s. = 180; s.l. = 450) N temp or cosmop temp
Veronicastrum (20) NE As, NE Am
Wulfenia (3) SE Eur, Turk
Wulfeniopsis (2) Afghan Nepal
Veronicaceae unassigned to tribe 3/53
Campylanthus (15) Macaron NE Af Arabia
Lafuentea (2) Spain
Morocco
Ourisia (36) Andes (20), NZ (15), Tasm (1)
Lithophytum (1) Mexico
Limnophila
(37) OW tropics
Peltanthera
(1) Costa Rica to Bolivia
Rehmannia
(9) E As
Sanango
(1) Ecuador Peru (Gesneriaceae?)
Wightia
(2) SE Asia epiphytes becoming trees
Excluded from Acanthaceae (possibly Schlegeliaceae)
Thomandersia
Baill. (6) trop Af
Excluded from Gesneriaceae (J. Smith, V. Savolainen p.c.)
Cubitanthus Barringer (1), Brazil
Acknowledgments:
P. Cantino (Lamiaceae)
E. Norman (Buddlejaceae,
Sanango, Peltanthera)
P. Garnock-Jones (New
Zealand Scrophulariaceae s.l.)
B. Oxelman (unpubl. Results
in Buddlejaceae, Scrophulariaceae s.l.)
J. Reveal (nomenclature
and misc.)
A. Weber (Gesneriaceae)
J. Smith (Gesneriaceae)
R. Harley (Lamiaceae)
P. Goldblatt
(Scrophulariaceae s.l.; esp. S. Africa)
D. Albach (Veroniceae and
Veronicaceae/Plantaginaceae)
Eva Wallander
(eva.wallander@systbot.gu.se) (Oleaceae)
R. Scotland (Acanthaceae)
B. Bremer (scroph tribe
Manuleeae, including former Selagineae)
H.-D. Ihlenfeldt
(Pedaliaceae, Martyniaceae, Trapellaceae)
D. Mabberley (Lamiaceae)
L. Lohmann (Bignoniaceae)
D. Nickrent
(Orobanchaceae)
D. Albach (Veronicaceae)