| Knowledge 
              of the timing and spatial structure of conifer invasion into meadows 
              is critical to understanding the natural dynamics of forest-meadow 
              mosaics and to establishing a baseline for assessing future change. In 
              this section, we present studies of the spatial and temporal patterns 
              of conifer invasion at Bunchgrass Ridge and of the extent to which 
              these have been shaped by extrinsic factors (changes in climate 
              or disturbance regime) and intrinsic controls (interactions within 
              and between tree species). A. 
              Spatial and temporal patternsB. Extrinsic vs. 
            intrinsic controls
 | 
               
                | Waves 
                  of conifer invasion |   
                | 
                     
                      |  |   
                      | 
                           
                            | • grand 
                              fir - live • lodgepole 
                              pine - live
 | • grand 
                              fir - dead • 
                              lodgepole pine - dead
 |   
                            | Symbol 
                              size approximates age |  |   
                      | 
                           
                            | Successive 
                              waves of conifer invasion in a 1-ha plot. Lodgepole 
                              pine (now dead) and grand fir were the first to 
                              invade open meadow in the mid- to late-1800s. More 
                              recent (20th-century) invasion of lodgepole 
                              pine (some dead) and grand fir has filled large 
                              areas of adjacent meadow. |  |  |  |